Castilleja attenuata |
Castilleja hololeuca |
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attenuate Indian paintbrush, attenuate paintbrush, narrow-leaf owl's-clover, narrow-leaf owl-clover, narrow-leaf paintbrush, valley-tassels |
Channel Islands paintbrush, island paintbrush, island white-felted paintbrush, white-felted paintbrush |
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Habit | Herbs, annual, 1–4.5 dm; with fibrous roots. | Shrubs or subshrubs, 3–10 dm; from a woody caudex; with thick, woody roots. |
Stems | solitary, erect to ascending, unbranched, sometimes branched from base, hairs spreading, short and medium, ± stiff, eglandular. |
many, erect to spreading, much-branched with many short axillary shoots, proximal stems reaching 1+ cm wide, white-felted, hairs dense, spreading to ± appressed, intertwined, long, soft, slightly branched, obscuring surface. |
Leaves | green to purple-tinged, linear to linear-lanceolate, (1–)2–8 cm, not fleshy, margins plane, flat, 0(–5)-lobed, apex acuminate; lobes spreading to ascending, filiform to lanceolate, apex sometimes acuminate. |
ash gray, leaves of previous year persisting on proximal stem, linear, older leaves sometimes broadly linear, 1–5 cm, not fleshy, margins plane, involute, 0-lobed, apex obtuse; lobes ascending, linear-lanceolate, apex obtuse to rounded, sometimes acute. |
Inflorescences | (1.5–)2–10(–19, –30 in fruit) × 1–2 cm; bracts proximally green to pale brown, rarely dull reddish brown, distally white on apices, sometimes pale yellow or pale pink-purplish on apices, rarely greenish or dull reddish brown throughout, lanceolate or lanceolate-elliptic, 3(–5)-lobed; lobes ascending, linear to lanceolate, long, arising below mid length, apex acuminate, acute, or obtuse. |
3–16 × 2–4 cm; bracts pale gray-green throughout, or proximally pale gray-green, distally red or yellow, proximal linear, distal broader, often oblanceolate to obovate, (0–)3-lobed; lobes spreading, linear to oblong, short to long, arising below mid length, apex rounded to truncate or acute. |
Corollas | straight, 10–25 mm; tube 9–20 mm, not expanded distally; beak exserted, straight, adaxially white, light yellow, or greenish, 3–5 mm, inconspicuously puberulent; abaxial lip white or yellow with deep brown to purple spots, often becoming pink, slightly inflated, exserted or not, pouches 3, 2 mm wide, 1–1.5 mm deep, 3–4 mm, 75–80% as long as beak; teeth erect, white, pale yellow, or pink, 0.5–1.2 mm. |
slightly curved in proximal 1/3, 14–26(–31) mm; tube 9–17 mm; abaxial lip not exserted, beak exserted; beak adaxially green to yellowish, 11–14 mm; abaxial lip ascending, deep green, reduced, 2–3 mm, less than 33% as long as beak; teeth reduced to apiculations, deep green, 1–1.5 mm. |
Calyces | colored as bracts, (8–)15–23 mm; abaxial and adaxial clefts (4–)6–8 mm, abaxial ca. 50% of calyx length, adaxial ca. 75% of calyx length, lateral 3–3.5 mm, 33% of calyx length; lobes linear to narrowly triangular, apex acute to acuminate. |
colored as bracts, with conspicuous whitish veins, 14–18(–20) mm; abaxial and adaxial clefts 5–12 mm, 30–67% of calyx length, deeper than laterals, lateral 0 mm or nearly so, ca. 0% of calyx length; lobes linear-lanceolate, apex rounded, truncate, or emarginate. |
Filaments | glabrous. |
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2n | = 24. |
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Castilleja attenuata |
Castilleja hololeuca |
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Phenology | Flowering (Feb–)Mar–Jun. | Flowering Mar–Oct. |
Habitat | Grasslands, pastures, moist margins of springs and streams, damp rocky slopes. | Coastal sage scrub, chaparral slopes and flats, ledges, forest edges. |
Elevation | 0–2100 m. (0–6900 ft.) | 0–500 m. (0–1600 ft.) |
Distribution |
AZ; CA; OR; WA; BC; Mexico (Baja California); South America (Chile)
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CA
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Discussion | Castilleja attenuata is a common and widespread species, ranging from southwestern Canada to northern Baja California, with several disjunct populations in central Chile. It is sensitive to competition from weeds. Disjunct populations in the Rincon Mountains in Pima County, Arizona, often have pink bracts but are otherwise typical. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja hololeuca is endemic to the four major islands of the northern Channel Islands. It is similar to C. lanata, a mostly Mexican species that reaches Arizona, New Mexico, and southwest Texas. However, C. hololeuca appears to be more closely related to C. foliolosa, a species of the California mainland. These three species, and C. galehintoniae G. L. Nesom of Nuevo León, all share a distinctive, thick indument of pale, matted, and often branched hairs on the stems and leaves. Large individuals of C. hololeuca form thick, woody trunks. On Anacapa Island, closest to the mainland, red-flowered forms of C. hololeuca predominate. On Santa Cruz Island, to the west, both red- and yellow-bracted forms are fairly common, usually in separate populations. Further offshore, on San Miguel and Santa Rosa islands, most plants are yellow to, occasionally, peach in coloration. Reports of this species from Santa Barbara and Santa Catalina Islands are referable to other species, mostly C. foliolosa. Populations of C. hololeuca historically declined from grazing by introduced game and livestock species. The plants are recovering well, following removal of the grazing animals, and are now common in many areas, especially on San Miguel Island. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 590. | FNA vol. 17, p. 614. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | Orthocarpus attenuatus | C. lanata subsp. hololeuca |
Name authority | (A. Gray) T. I. Chuang & Heckard: Syst. Bot. 16: 656. (1991) | Greene: W. Amer. Sci. 3: 3. (1886) — (as Castilleia) |
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