Castilleja angustifolia |
Castilleja linariifolia |
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narrow-leaf paintbrush, northwestern Indian paintbrush, northwestern paintbrush, violet desert paintbrush |
desert paintbrush, desert paintbursh, linaria-leaf Indian paintbrush, narrow leaf paintbrush, narrow-leaf Indian paintbrush, Wyoming Indian paintbrush, Wyoming or narrow-leaf or long-leaf paintbrush, Wyoming paintbrush |
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Habit | Herbs, perennial, 0.9–3.8(–4) dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.8–10(–20) dm; from a woody caudex; with a taproot or branching roots. | ||||||||
Stems | few to many, ascending to erect, branched, especially near base, sometimes unbranched, hairs sparse to dense, spreading to retrorse, long, sometimes short, soft to stiff, usually mixed with short-glandular ones, sometimes viscid. |
few to many, ascending to erect, much-branched, glabrous proximally, hairy distally, sometimes glabrous or hairy throughout, hairs sparse, sometimes dense, spreading to slightly retrorse, short to long, ± stiff, eglandular. |
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Leaves | brown or purplish, sometimes green, linear to lanceolate or broadly lanceolate, 1.2–7(–7.5) cm, not fleshy, margins plane, sometimes ± wavy, involute or flat, (0–)3–5-lobed, rarely with secondary lobes, apex acuminate to rounded; lobes spreading, oblong or lanceolate to linear-lanceolate, apex acute to rounded. |
± yellow to gray-green, sometimes becoming ± purple, linear to narrowly lanceolate, (1–)2–10 cm, not fleshy, rarely ± thickened and fleshy, margins plane, involute, 0–3(–5)-lobed, apex acute or acuminate; lobes spreading, linear, apex acute to acuminate. |
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Inflorescences | 2.5–20 × 1.5–5 cm; bracts proximally greenish or dull purplish, distally pink, magenta, pink-purple, reddish pink, pale yellow, pale yellow-orange, pale orange, or white, rarely reddish or orange-red, lanceolate to oblong, 3–5(–9)-lobed, sometimes with secondary lobes; lobes spreading or ascending, oblanceolate or linear, proximal lobes often much longer than distal, proximal lobes arising below or a little above mid length, apex acute to rounded. |
4–20 × 2–7.5 cm; bracts red to red-orange throughout, sometimes pale green, yellow, magenta, pink-purple, or white throughout, or proximally pale greenish to straw colored, distally colored as above, lanceolate to narrowly lanceolate or narrowly oblong, 3(–5)-lobed; lobes spreading to ascending, narrowly lanceolate to linear, shorter than central lobe, arising in proximal 1/3, apex obtuse to acuminate. |
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Corollas | straight, 18–27(–32) mm; tube 8–17 mm; beak usually long-exserted, adaxially green or pink, 8–15 mm; abaxial lip deep green, reduced, inconspicuous, 1–2.5 mm, 5–20% as long as beak; teeth incurved to ascending, deep green, 0.5–1.5 mm. |
often slightly curved, 25–45 mm; tube 11–22(–25) mm; beak exserted, longer than calyx lobes, usually projecting through abaxial cleft; beak adaxially yellow-green or green, 9–21(–24) mm; abaxial lip deep green, reduced, often visible in exserted sideways through abaxial calyx cleft, 0.5–3 mm, 10–15% as long as beak; teeth incurved, green or whitish, 5–2(–3) mm. |
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Calyces | proximally green, yellow, brown, or purple, lobes colored as bract lobes, sometimes with a yellow band between proximal and distal parts, 13–25(–28) mm; abaxial clefts 3–8 mm, adaxial 5–9(–12) mm, clefts 30–50% of calyx length, deeper than laterals, lateral (1–)1.5–4(–5) mm, 10–25% of calyx length; lobes lanceolate to oblong, abaxials wider than adaxials, apex acute to rounded. |
proximally greenish, whitish, or yellowish, distally colored as bracts, 18–30(–35) mm; abaxial clefts 10–20(–22) mm, adaxial 2–6(–12) mm, abaxial ca. 70% of calyx length, adaxial ca. 20–25% of calyx length, deeper than laterals, lateral 1.5–5(–6) mm, 12–17% of calyx length; lobes curved slightly toward adaxial side, narrowly oblong to narrowly lanceolate, apex acute. |
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2n | = 24, 48. |
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Castilleja angustifolia |
Castilleja linariifolia |
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Phenology | Flowering Apr–Oct. | |||||||||
Habitat | Sagebrush steppes, grasslands, dry rocky slopes and flats, open forests, talus, lowlands to montane, occasionally subalpine. | |||||||||
Elevation | 600–3400 m. (2000–11200 ft.) | |||||||||
Distribution |
ID; MT; NV; OR; SD; UT; WY
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AZ; CA; CO; ID; MT; NM; NV; OR; UT; WY
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Discussion | Varieties 3 (3 in the flora). Much confusion exists concerning Castilleja angustifolia and the closely related C. chromosa. Sometimes C. chromosa is treated as a variety of C. angustifolia, using the name C. angustifolia var. dubia. The latter name is used here to represent a different assemblage of plants, not including C. chromosa. At other times, C. chromosa is synonymized completely under C. angustifolia. However, the two species are in most cases easily separable, and where they are sympatric there is little evidence of intergradation. Both C. angustifolia var. dubia and C. chromosa are accepted here. See additional comments under 3b. C. angustifolia var. dubia and 15. C. chromosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja linariifolia is widespread in the western United States and is important to Native Americans as a source of dyes and for medicinal and ceremonial purposes (D. E. Moerman 1998). It is the state flower of Wyoming. Castilleja linariifolia is closely associated with, and undoubtedly parasitic on, the roots of sagebrush, Artemisia tridentata. The bracts are usually red to red-orange, with uncommon yellow-bracted variants. On the eastern slope of the Sierra Nevada in California, most populations have pink-purple to magenta bracts, and at least one population has almost pure white bracts. Plants associated with hot springs in the Ash Meadows area of Nye County, Nevada, are unusually tall and have somewhat fleshy leaves. Castilleja linariifolia stems are usually glabrous, but pubescent stems are sporadic across its range. However, in the Mt. Charleston area of the Spring Mountains, in southern Nevada, most plants are pubescent, ranging from an inconspicuous layer to a fairly dense and obvious indument. These plants have been called forma omnipubescens Pennell. Elsewhere, especially in central and northern Arizona, individuals with short-pubescent stems are intermingled with more typical plants. In addition, late-blooming forms associated with hot spring sites in the eastern Mojave Desert are particularly thick-stemmed and vigorous. Hybrids between C. linariifolia and C. scabrida are known from Garfield County, Utah. Castilleja linariifolia is parapatric with the similar C. wootonii in central New Mexico. Castilleja linariifolia is reported to hybridize with C. chromosa, C. flava, and C. miniata. A possible hybrid with C. septentrionalis was named C. ×cognata Greene. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 586. | FNA vol. 17, p. 622. | ||||||||
Parent taxa | ||||||||||
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Synonyms | Euchroma angustifolia | C. trainii | ||||||||
Name authority | (Nuttall) G. Don: Gen. Hist. 4: 616. 1837/1838 | Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 532. (1846) — (as lineariaefolia) | ||||||||
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