Castilleja angustifolia |
Castilleja gleasoni |
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narrow-leaf paintbrush, northwestern Indian paintbrush, northwestern paintbrush, violet desert paintbrush |
Mount Gleason paintbrush, Mt. Gleason paintbrush |
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Habit | Herbs, perennial, 0.9–3.8(–4) dm; from a woody caudex; with a taproot. | Herbs or subshrubs, perennial, 3–8 dm; from a woody caudex; with a taproot. | ||||||||
Stems | few to many, ascending to erect, branched, especially near base, sometimes unbranched, hairs sparse to dense, spreading to retrorse, long, sometimes short, soft to stiff, usually mixed with short-glandular ones, sometimes viscid. |
several to many, erect to ascending, branched, sometimes unbranched, hairs moderately dense, spreading, ± matted, ash gray, branched, sometimes unbranched, medium length, soft, often mixed with shorter, unbranched, stipitate-glandular ones, not obscuring surface. |
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Leaves | brown or purplish, sometimes green, linear to lanceolate or broadly lanceolate, 1.2–7(–7.5) cm, not fleshy, margins plane, sometimes ± wavy, involute or flat, (0–)3–5-lobed, rarely with secondary lobes, apex acuminate to rounded; lobes spreading, oblong or lanceolate to linear-lanceolate, apex acute to rounded. |
ash gray, linear-lanceolate to narrowly oblong or narrowly oblanceolate, 2–6 cm, not fleshy, margins wavy, involute, 0(–3)-lobed, apex acute to obtuse; lobes spreading, narrowly lanceolate, apex acute to obtuse. |
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Inflorescences | 2.5–20 × 1.5–5 cm; bracts proximally greenish or dull purplish, distally pink, magenta, pink-purple, reddish pink, pale yellow, pale yellow-orange, pale orange, or white, rarely reddish or orange-red, lanceolate to oblong, 3–5(–9)-lobed, sometimes with secondary lobes; lobes spreading or ascending, oblanceolate or linear, proximal lobes often much longer than distal, proximal lobes arising below or a little above mid length, apex acute to rounded. |
10–15(–30) cm; bracts proximally greenish to dull brownish purple, distally red to deep red or red-orange, lanceolate to oblong or narrowly ovate, (0–)3(–5)-lobed; lobes ascending to spreading, narrowly oblong to narrowly lanceolate, medium length, arising below mid length, center lobe apex obtuse or toothed, lateral ones acute. |
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Corollas | straight, 18–27(–32) mm; tube 8–17 mm; beak usually long-exserted, adaxially green or pink, 8–15 mm; abaxial lip deep green, reduced, inconspicuous, 1–2.5 mm, 5–20% as long as beak; teeth incurved to ascending, deep green, 0.5–1.5 mm. |
± straight, 18–30 mm; tube 9–15 mm; beak exserted, adaxially yellow, 9–15(–20) mm; abaxial lip spreading, deep green to ± black, reduced, 0.5–2 mm, 20% as long as beak; teeth incurved, green, 0.2–1 mm. |
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Calyces | proximally green, yellow, brown, or purple, lobes colored as bract lobes, sometimes with a yellow band between proximal and distal parts, 13–25(–28) mm; abaxial clefts 3–8 mm, adaxial 5–9(–12) mm, clefts 30–50% of calyx length, deeper than laterals, lateral (1–)1.5–4(–5) mm, 10–25% of calyx length; lobes lanceolate to oblong, abaxials wider than adaxials, apex acute to rounded. |
proximally pale to green, sometimes dull purple, often paler or greener than bracts, distal 1/4 or less colored as bract lobes, 12–17 mm; abaxial and adaxial clefts 3–8 mm, 40% of calyx length, deeper than laterals, lateral 0.5–3 mm, 20–25% of calyx length; lobes ovate to narrowly triangular, apex obtuse to acute. |
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2n | = 72. |
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Castilleja angustifolia |
Castilleja gleasoni |
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Phenology | Flowering May–Jun. | |||||||||
Habitat | Ledges, rocky slopes, open yellow pine forests, montane chaparral or sagebrush. | |||||||||
Elevation | 900–2200 m. (3000–7200 ft.) | |||||||||
Distribution |
ID; MT; NV; OR; SD; UT; WY
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CA |
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Discussion | Varieties 3 (3 in the flora). Much confusion exists concerning Castilleja angustifolia and the closely related C. chromosa. Sometimes C. chromosa is treated as a variety of C. angustifolia, using the name C. angustifolia var. dubia. The latter name is used here to represent a different assemblage of plants, not including C. chromosa. At other times, C. chromosa is synonymized completely under C. angustifolia. However, the two species are in most cases easily separable, and where they are sympatric there is little evidence of intergradation. Both C. angustifolia var. dubia and C. chromosa are accepted here. See additional comments under 3b. C. angustifolia var. dubia and 15. C. chromosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja gleasoni is an uncommon plant endemic to the upper elevations of the San Gabriel Mountains of Los Angeles County. It was treated as a polyploid derivative of C. affinis var. affinis (as subsp. affinis) × C. foliolosa (T. I. Chuang and L. R. Heckard 1993b), a hypothetical ancestry supported by a chromosome number of 2n = 72. Others have placed it as a subspecies of C. pruinosa, a similar species. However, the morphology of C. gleasoni also suggests it could have originated as a cross between the diploid C. foliolosa and a tetraploid form of C. martini; careful morphological and molecular analyses are needed to determine its true ancestry. Castilleja gleasoni is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 586. | FNA vol. 17, p. 610. | ||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Euchroma angustifolia | C. pruinosa subsp. gleasoni | ||||||||
Name authority | (Nuttall) G. Don: Gen. Hist. 4: 616. 1837/1838 | Elmer: Bot. Gaz. 39: 51. (1905) — (as Castilleia) | ||||||||
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