Castilleja angustifolia |
Castilleja cinerea |
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narrow-leaf paintbrush, northwestern Indian paintbrush, northwestern paintbrush, violet desert paintbrush |
ash-gray Indian paintbrush, ash-gray paintbrush |
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Habit | Herbs, perennial, 0.9–3.8(–4) dm; from a woody caudex; with a taproot. | Herbs, perennial, 0.5–3 dm; from a woody caudex; with a taproot. | ||||||||
Stems | few to many, ascending to erect, branched, especially near base, sometimes unbranched, hairs sparse to dense, spreading to retrorse, long, sometimes short, soft to stiff, usually mixed with short-glandular ones, sometimes viscid. |
several to many, erect to ascending, or decumbent, inflorescence erect in high-elevation form, unbranched, sometimes branched, hairs dense, spreading, ashy gray, short and long, soft, mixed with short stipitate-glandular ones. |
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Leaves | brown or purplish, sometimes green, linear to lanceolate or broadly lanceolate, 1.2–7(–7.5) cm, not fleshy, margins plane, sometimes ± wavy, involute or flat, (0–)3–5-lobed, rarely with secondary lobes, apex acuminate to rounded; lobes spreading, oblong or lanceolate to linear-lanceolate, apex acute to rounded. |
green, brown, purple, or deep gray, linear or narrowly to broadly lanceolate to sometimes ovate, 0.7–3 cm, not fleshy, margins plane, slightly involute, 0(–3)-lobed, apex acuminate; lobes ascending to spreading, linear to lanceolate, apex acuminate. |
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Inflorescences | 2.5–20 × 1.5–5 cm; bracts proximally greenish or dull purplish, distally pink, magenta, pink-purple, reddish pink, pale yellow, pale yellow-orange, pale orange, or white, rarely reddish or orange-red, lanceolate to oblong, 3–5(–9)-lobed, sometimes with secondary lobes; lobes spreading or ascending, oblanceolate or linear, proximal lobes often much longer than distal, proximal lobes arising below or a little above mid length, apex acute to rounded. |
1–8.5 × 2–5 cm; bracts proximally greenish or deep reddish purple, distally burnt orange, sometimes yellow or deep red to deep burgundy, proximal sometimes lanceolate with narrow lobes, distal or all bracts broadly lanceolate to oblong or slightly oblanceolate, (0–)3–5-lobed, appearing dusty with dense, short stipitate-glandular hairs, many with a nodulose to pillarlike, crystallized, usually pigmented exudate, papillose at 40x; lobes ascending-spreading, oblong or oblanceolate, short, arising above mid length, central lobe apex rounded, often expanded, rounded, or truncate, lateral ones acute to rounded. |
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Corollas | straight, 18–27(–32) mm; tube 8–17 mm; beak usually long-exserted, adaxially green or pink, 8–15 mm; abaxial lip deep green, reduced, inconspicuous, 1–2.5 mm, 5–20% as long as beak; teeth incurved to ascending, deep green, 0.5–1.5 mm. |
straight, 12–18 mm; tube 9–14 mm; beak included or tip just barely exserted, adaxially green or pale yellow to deep burgundy, 3–5 mm; abaxial lip green, burgundy, or reddish purple (in high-elevation form), little inflated, small, included, 2 mm, to 20% as long as beak; teeth incurved, green, 0.2–0.5 mm. |
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Calyces | proximally green, yellow, brown, or purple, lobes colored as bract lobes, sometimes with a yellow band between proximal and distal parts, 13–25(–28) mm; abaxial clefts 3–8 mm, adaxial 5–9(–12) mm, clefts 30–50% of calyx length, deeper than laterals, lateral (1–)1.5–4(–5) mm, 10–25% of calyx length; lobes lanceolate to oblong, abaxials wider than adaxials, apex acute to rounded. |
colored as bracts, sometimes whitish proximally, 1.5–20 mm (shorter in upper elevation form); abaxial and adaxial clefts 3.5–8 mm, 30–50% of calyx length, all 4 clefts subequal; lobes linear to narrowly oblong or oblanceolate, apex obtuse to rounded, densely stipitate-glandular. |
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Castilleja angustifolia |
Castilleja cinerea |
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Phenology | Flowering May–Aug(–Oct). | |||||||||
Habitat | Dry rocky slopes, ridges, and flats, pebble plains, sagebrush openings, open conifer forests. | |||||||||
Elevation | 1800–3100 m. [5900–10200 ft.] | |||||||||
Distribution |
ID; MT; NV; OR; SD; UT; WY
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CA
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Discussion | Varieties 3 (3 in the flora). Much confusion exists concerning Castilleja angustifolia and the closely related C. chromosa. Sometimes C. chromosa is treated as a variety of C. angustifolia, using the name C. angustifolia var. dubia. The latter name is used here to represent a different assemblage of plants, not including C. chromosa. At other times, C. chromosa is synonymized completely under C. angustifolia. However, the two species are in most cases easily separable, and where they are sympatric there is little evidence of intergradation. Both C. angustifolia var. dubia and C. chromosa are accepted here. See additional comments under 3b. C. angustifolia var. dubia and 15. C. chromosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja cinerea is endemic to the higher elevations of the San Bernardino Mountains, San Bernardino County. Most plants are upright to ascending and have yellow to yellow-orange inflorescences, with occasional plants ranging to dull red, especially with age. On Sugarloaf Mountain, mostly above 2700 m, is a distinctive form with consistently reddish purple to burgundy inflorescences and a strongly decumbent growth form. Castilleja cinerea is most often associated with and likely parasitic on Artemisia nova and Eriogonum species. Castilleja cinerea is known from few populations and is threatened by livestock grazing, development, and vehicle use. It is listed as a threatened species under the Endangered Species Act of the United States. The crystalline exudate associated with the stipitate-glandular pubescence of the distal portion of the bracts is unique in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 586. | FNA vol. 17, p. 597. | ||||||||
Parent taxa | ||||||||||
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Synonyms | Euchroma angustifolia | Orthocarpus cinereus | ||||||||
Name authority | (Nuttall) G. Don: Gen. Hist. 4: 616. 1837/1838 | A. Gray: Proc. Amer. Acad. Arts 19: 93. (1883) — (as Castilleia) | ||||||||
Web links |