Castilleja affinis |
Castilleja sessiliflora |
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California paintbrush, coast Indian paintbrush, coast paintbrush, Indian paintbrush |
downy paintedcup, Great Plains Indian paintbrush |
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Habit | Herbs, perennial, 1.4–6 dm; caudex woody; with a taproot. | Herbs, perennial, 1–4 dm; from a branching, woody caudex; with a taproot. | ||||||||
Stems | few to many, erect to ascending, unbranched or branched at proximal nodes, sometimes with short, leafy axillary branches, hairs sparse to dense, spreading, short and long, soft to stiff, unbranched, sometimes branched, eglandular, sometimes stipitate-glandular. |
few to many, ascending to erect, often decumbent at base, unbranched, sometimes branched, hairs spreading, sometimes matted, short to medium length, ± soft, eglandular, often with a layer of minute-glandular hairs, sometimes woolly. |
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Leaves | green or purplish, sometimes red-brown, linear or linear-oblong to broadly lanceolate, 2–13 cm, not or ± fleshy, margins wavy or plane, flat or involute, 0–5-lobed, sometimes with small secondary lobes, short, leafy axillary shoots common, usually conspicuous, apex acuminate or acute to rounded; lobes spreading, linear to lanceolate, lateral lobes almost as wide as mid blade, apex acute to rounded. |
green to purple, or grayish with dust and hairs, linear to narrowly lanceolate, (1–)2–5(–6) cm, not fleshy, margins plane, involute, (0–)3–5-lobed, apex acuminate to acute; lobes divergent, spreading, linear, apex acute. |
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Inflorescences | 3–25(–30) × 1.5–5 cm; bracts proximally green or deep purple, distally red, crimson, scarlet, pink, pinkish purple, pinkish red, or yellow, sometimes rose magenta, red-orange, or orange, oblanceolate or obovate to oblong or lanceolate, (0–)3–5(–7)-lobed; lobes spreading to ascending, linear to obovate, long, proximal lobes at or arising below mid length, apices acute to obtuse, center lobe sometimes rounded. |
3–18 × 2.5–6.5 cm; bracts green to purplish throughout, sometimes reddish brown, pink, or lavender throughout, or distally white or pale yellow, sometimes distally dull pink, pink, salmon, orangish, pale pink-orange, buff, or cream, lanceolate, similar to distal leaves, 3(–5)-lobed; lobes spreading, linear-lanceolate, long, arising at or below mid length, apex acute to acuminate, sometimes obtuse. |
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Pedicels | 0 mm or nearly so. |
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Corollas | straight or ± curved, 17–40 mm; tube 10–15 mm; beak long-exserted to subequal to calyx, adaxially green, 7–20 mm, puberulent, eglandular; abaxial lip deep green to reddish, brown, or deep purple, reduced, inconspicuous, included in calyx, 1.5–3 mm, 15–25% as long as beak; teeth ascending, green, 0.5–2 mm. |
strongly curved distally, 35–55 mm; tube 24–45 mm; abaxial lip, beak, and distal portion of tube exserted; beak adaxially green, yellow, pinkish, purplish, or whitish, 9–15 mm; abaxial lip green, pale green, or purple, protruding, shelflike, 4–8 mm, 50–70% as long as beak; teeth spreading, white, pale yellow, pink, or purple, 3–4 mm. |
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Calyces | proximally pale or green, distally as in distal portion of bracts, 14–35 mm; abaxial and adaxial clefts 6–22 mm, 33–50% of calyx length, deeper than laterals, lateral 2–7 mm, 15–25% of calyx length; lobes lanceolate or oblong, apex acute to obtuse or rounded, sometimes curved upward. |
colored as bracts, sometimes proximally white, 20–40 mm; abaxial and adaxial clefts 12–20 mm, 40–60% of calyx length, deeper than laterals, lateral 5–15 mm, 10–25% of calyx length; lobes linear, apex acute to acuminate. |
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2n | = 48, 72, 96. |
= 24. |
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Castilleja affinis |
Castilleja sessiliflora |
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Phenology | Flowering (Feb–)Mar–Aug(–Oct). | |||||||||
Habitat | Dry mixed grass and shortgrass prairies, prairie sandhills, sandsage plains, sand prairies, rocky or sandy slopes, bluffs, open forests, or desert scrub, limestone, sandstone, gypsum, granite, other bedrock types. | |||||||||
Elevation | 0–2300 m. (0–7500 ft.) | |||||||||
Distribution |
CA; nw Mexico
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AZ; CO; IA; IL; KS; MN; MO; MT; ND; NE; NM; OK; SD; TX; WI; WY; AB; MB; SK; Mexico (Coahuila, Nuevo León, Tamaulipas)
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Discussion | Varieties 3 (3 in the flora). Castilleja affinis is highly variable and one of the more common paintbrushes at lower elevations in California, west of the Sierra Nevada, from the northern coast south to northern Baja California. Some recent authors (for example, M. Wetherwax et al. 2012) include C. litoralis of the Pacific Northwest coast as a subspecies of C. affinis, but due to the high polyploid nature of C. litoralis and its significantly closer morphological resemblance to C. miniata var. dixonii, that treatment is not followed here. Also see the comments under 62. C. litoralis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja sessiliflora ranges across the Great Plains from southern Canada to northern Mexico, where it is apparently rare. In Texas and northern Mexico, its range overlaps with the similar C. mexicana. Most populations of C. sessiliflora, especially north of Texas, have white to pale yellow inflorescences; in southwestern Texas they are more variable in color, with pink-purple plants often predominating locally. Those plants with pink-purple inflorescences were named forma purpurina by F. W. Pennell. In the limestone deserts of southern New Mexico and southeastern Arizona, the inflorescences are often a pale pink-orange, but these are intermingled with more typical greenish white plants. Occasional hybrids between C. angustifolia var. dubia and C. sessiliflora are known from northeastern Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 582. | FNA vol. 17, p. 657. | ||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | Hooker & Arnott: Bot. Beechey Voy., 154. (1833) | Pursh: Fl. Amer. Sept. 2: 738. (1813) | ||||||||
Web links |