Castilleja affinis |
Castilleja peckiana |
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California paintbrush, coast Indian paintbrush, coast paintbrush, Indian paintbrush |
peak paintbrush, Peck's Indian paintbrush, Peck's paintbrush |
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Habit | Herbs, perennial, 1.4–6 dm; caudex woody; with a taproot. | Herbs, perennial, (1.8–)2.4–6 dm; from a woody caudex; with a stout taproot. | ||||||||
Stems | few to many, erect to ascending, unbranched or branched at proximal nodes, sometimes with short, leafy axillary branches, hairs sparse to dense, spreading, short and long, soft to stiff, unbranched, sometimes branched, eglandular, sometimes stipitate-glandular. |
few to many, erect or ascending, often branched distally, sometimes unbranched, sometimes with short axillary shoots, proximal hairs retrorse to appressed, short to moderately long, distal hairs spreading, longer, soft, often mixed with short stipitate-glandular ones. |
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Leaves | green or purplish, sometimes red-brown, linear or linear-oblong to broadly lanceolate, 2–13 cm, not or ± fleshy, margins wavy or plane, flat or involute, 0–5-lobed, sometimes with small secondary lobes, short, leafy axillary shoots common, usually conspicuous, apex acuminate or acute to rounded; lobes spreading, linear to lanceolate, lateral lobes almost as wide as mid blade, apex acute to rounded. |
green, linear-lanceolate, rarely broadly lanceolate, (1.2–)2.5–8(–9) cm, not fleshy, margins plane, flat or involute, 0(–3)-lobed, apex acute to rounded; lobes ascending-spreading, narrowly lanceolate to linear, apex acute or obtuse. |
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Inflorescences | 3–25(–30) × 1.5–5 cm; bracts proximally green or deep purple, distally red, crimson, scarlet, pink, pinkish purple, pinkish red, or yellow, sometimes rose magenta, red-orange, or orange, oblanceolate or obovate to oblong or lanceolate, (0–)3–5(–7)-lobed; lobes spreading to ascending, linear to obovate, long, proximal lobes at or arising below mid length, apices acute to obtuse, center lobe sometimes rounded. |
(2–)4–17 × 1.5–3 cm; bracts proximally greenish, distally red, orange-red, or orange, proximal sometimes lanceolate, distal broadly lanceolate to ovate, (0–)3(–7)-lobed; lobes spreading to ascending, linear to lanceolate, long, arising near or below mid length, central lobe apex rounded, lateral ones acute. |
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Pedicels | 0 mm or nearly so. |
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Corollas | straight or ± curved, 17–40 mm; tube 10–15 mm; beak long-exserted to subequal to calyx, adaxially green, 7–20 mm, puberulent, eglandular; abaxial lip deep green to reddish, brown, or deep purple, reduced, inconspicuous, included in calyx, 1.5–3 mm, 15–25% as long as beak; teeth ascending, green, 0.5–2 mm. |
straight, 23–30(–35) mm; tube 12–20 mm; beak exserted, adaxially green, 8–12(–14) mm; abaxial lip pale to deep green, reduced, rounded, 0.5–1.5 mm, 10–20% as long as beak; teeth erect to incurved, green, (0.4–)0.7–1.2(–2) mm. |
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Calyces | proximally pale or green, distally as in distal portion of bracts, 14–35 mm; abaxial and adaxial clefts 6–22 mm, 33–50% of calyx length, deeper than laterals, lateral 2–7 mm, 15–25% of calyx length; lobes lanceolate or oblong, apex acute to obtuse or rounded, sometimes curved upward. |
proximally pale yellow or greenish, distally colored as bracts, (15–)18–28 mm; abaxial and adaxial clefts 6–12 mm, 40–45% of calyx length, deeper than laterals, lateral 2.5–8 mm, 20–30% of calyx length; lobes linear to narrowly lanceolate, apex acute to acuminate. |
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2n | = 48, 72, 96. |
= 72, 96, ca. 120. |
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Castilleja affinis |
Castilleja peckiana |
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Phenology | Flowering (Apr–)May–Aug. | |||||||||
Habitat | Open conifer forests, sagebrush slopes, riparian meadows, shores. | |||||||||
Elevation | 1400–2600 m. (4600–8500 ft.) | |||||||||
Distribution |
CA; nw Mexico
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ID; NV; OR
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Discussion | Varieties 3 (3 in the flora). Castilleja affinis is highly variable and one of the more common paintbrushes at lower elevations in California, west of the Sierra Nevada, from the northern coast south to northern Baja California. Some recent authors (for example, M. Wetherwax et al. 2012) include C. litoralis of the Pacific Northwest coast as a subspecies of C. affinis, but due to the high polyploid nature of C. litoralis and its significantly closer morphological resemblance to C. miniata var. dixonii, that treatment is not followed here. Also see the comments under 62. C. litoralis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
When describing Castilleja peckiana, Pennell noted that its variation approached C. hispida on one hand and C. miniata on the other, and it is likely of allopolyploid derivation. L. R. Heckard (1968) found chromosome numbers of n = 36, 48, and ca. 60. He hypothesized that C. chromosa, C. hispida var. acuta, and C. miniata were likely involved in its ancestry, and possibly C. pruinosa as well. Heckard suggested subsequent introgression among the derived forms introduced further complexity. Though complex, these forms are self-perpetuating and appear morphologically stable within their range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 582. | FNA vol. 17, p. 643. | ||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||
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Subordinate taxa | ||||||||||
Name authority | Hooker & Arnott: Bot. Beechey Voy., 154. (1833) | Pennell: Notul. Nat. Acad. Nat. Sci. Philadelphia 74: 9. (1941) | ||||||||
Web links |