Castilleja affinis |
Castilleja occidentalis |
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California paintbrush, coast Indian paintbrush, coast paintbrush, Indian paintbrush |
western Indian paintbrush, western or western yellow paintbrush, western paintbrush |
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Habit | Herbs, perennial, 1.4–6 dm; caudex woody; with a taproot. | Herbs, perennial, 0.7–2(–3) dm; from a small, woody caudex; with a taproot. | ||||||||
Stems | few to many, erect to ascending, unbranched or branched at proximal nodes, sometimes with short, leafy axillary branches, hairs sparse to dense, spreading, short and long, soft to stiff, unbranched, sometimes branched, eglandular, sometimes stipitate-glandular. |
several to many, erect or ascending, usually short-decumbent at base, unbranched, sometimes glabrous proximally, hairs spreading, long, soft, mixed with medium length to short stipitate-glandular ones only in inflorescence. |
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Leaves | green or purplish, sometimes red-brown, linear or linear-oblong to broadly lanceolate, 2–13 cm, not or ± fleshy, margins wavy or plane, flat or involute, 0–5-lobed, sometimes with small secondary lobes, short, leafy axillary shoots common, usually conspicuous, apex acuminate or acute to rounded; lobes spreading, linear to lanceolate, lateral lobes almost as wide as mid blade, apex acute to rounded. |
green to deep purple, linear-lanceolate to broadly lanceolate (to linear on sterile shoots), 1.5–4(–5.5) cm, not fleshy, margins plane, flat, prominently veined, 0–3(–5)-lobed, apex acute to rounded; lobes ascending, lanceolate, apex acute. |
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Inflorescences | 3–25(–30) × 1.5–5 cm; bracts proximally green or deep purple, distally red, crimson, scarlet, pink, pinkish purple, pinkish red, or yellow, sometimes rose magenta, red-orange, or orange, oblanceolate or obovate to oblong or lanceolate, (0–)3–5(–7)-lobed; lobes spreading to ascending, linear to obovate, long, proximal lobes at or arising below mid length, apices acute to obtuse, center lobe sometimes rounded. |
2–7(–10) × 1–3.5 cm; bracts greenish to pale greenish yellow throughout, often aging dull reddish brown or reddish purple proximally, rarely dull reddish brown throughout, or proximally green, dull reddish brown, or reddish purple, distally greenish white, yellow, or cream, broadly lanceolate to widely oblong to ovate, 0–3(–7)-lobed; lobes ascending, triangular to lanceolate, medium length, usually arising at or above mid length, rarely just below, central lobe apex obtuse to rounded, others acute. |
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Pedicels | 0 mm or nearly so. |
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Corollas | straight or ± curved, 17–40 mm; tube 10–15 mm; beak long-exserted to subequal to calyx, adaxially green, 7–20 mm, puberulent, eglandular; abaxial lip deep green to reddish, brown, or deep purple, reduced, inconspicuous, included in calyx, 1.5–3 mm, 15–25% as long as beak; teeth ascending, green, 0.5–2 mm. |
straight, 16–25 mm; tube 9–15 mm; teeth and part of abaxial lip sometimes exserted, beak exserted; beak adaxially green, (2.5–)5–9 mm; abaxial lip green, reduced, often visible through abaxial cleft, slightly pouched, 1.5–3 mm, 25–50% as long as beak; teeth incurved to ascending, white, sometimes green, 0.7–2 mm. |
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Calyces | proximally pale or green, distally as in distal portion of bracts, 14–35 mm; abaxial and adaxial clefts 6–22 mm, 33–50% of calyx length, deeper than laterals, lateral 2–7 mm, 15–25% of calyx length; lobes lanceolate or oblong, apex acute to obtuse or rounded, sometimes curved upward. |
proximally green to purple, distally colored as bracts, 12–20 mm; abaxial and adaxial clefts 5–9(–10) mm, 40–50% of calyx length, deeper than laterals, lateral 1–3(–4.5) mm, 5–20% of calyx length; lobes lanceolate to triangular, apex acute, obtuse, or rounded. |
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2n | = 48, 72, 96. |
= 24, 48. |
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Castilleja affinis |
Castilleja occidentalis |
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Phenology | Flowering Jun–Sep. | |||||||||
Habitat | Meadows, gravel slopes, talus, ridges, mostly upper subalpine to alpine. | |||||||||
Elevation | 1500–4300 m. (4900–14100 ft.) | |||||||||
Distribution |
CA; nw Mexico
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CO; MT; NM; UT; AB; BC
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Discussion | Varieties 3 (3 in the flora). Castilleja affinis is highly variable and one of the more common paintbrushes at lower elevations in California, west of the Sierra Nevada, from the northern coast south to northern Baja California. Some recent authors (for example, M. Wetherwax et al. 2012) include C. litoralis of the Pacific Northwest coast as a subspecies of C. affinis, but due to the high polyploid nature of C. litoralis and its significantly closer morphological resemblance to C. miniata var. dixonii, that treatment is not followed here. Also see the comments under 62. C. litoralis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja occidentalis has a wide distribution in the southern Rocky Mountains, a gap in its distribution in Wyoming, and reappears in Montana and the Canadian Rockies. Bract lobing and color vary considerably but without correlation to geography. Although C. occidentalis resembles an alpine form of C. septentrionalis, C. occidentalis is missing in several regions in the distribution of the latter, even when extensive areas of suitable habitat are available, and has a discrete range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 582. | FNA vol. 17, p. 635. | ||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||
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Subordinate taxa | ||||||||||
Name authority | Hooker & Arnott: Bot. Beechey Voy., 154. (1833) | Torrey: Ann. Lyceum Nat. Hist. New York 2: 230. (1827) | ||||||||
Web links |