Castilleja affinis |
Castilleja lasiorhyncha |
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California paintbrush, coast Indian paintbrush, coast paintbrush, Indian paintbrush |
San Bernardino Mountains Indian paintbrush, San Bernardino Mountains paintbrush, San Bernardino Mountains. owl's clover |
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Habit | Herbs, perennial, 1.4–6 dm; caudex woody; with a taproot. | Herbs, annual, (0.6–)1–3(–4) dm; with fibrous roots. | ||||||||
Stems | few to many, erect to ascending, unbranched or branched at proximal nodes, sometimes with short, leafy axillary branches, hairs sparse to dense, spreading, short and long, soft to stiff, unbranched, sometimes branched, eglandular, sometimes stipitate-glandular. |
solitary, erect, unbranched or branched, hairs spreading, medium length and long, soft to ± stiff, mixed with shorter stipitate-glandular ones. |
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Leaves | green or purplish, sometimes red-brown, linear or linear-oblong to broadly lanceolate, 2–13 cm, not or ± fleshy, margins wavy or plane, flat or involute, 0–5-lobed, sometimes with small secondary lobes, short, leafy axillary shoots common, usually conspicuous, apex acuminate or acute to rounded; lobes spreading, linear to lanceolate, lateral lobes almost as wide as mid blade, apex acute to rounded. |
green to purple, linear to narrowly lanceolate, 0.5–4.2 cm, not fleshy, margins plane, flat, 0(–5)-lobed, apex acuminate to acute; lobes ascending to erect, linear to narrowly lanceolate, apex acuminate. |
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Inflorescences | 3–25(–30) × 1.5–5 cm; bracts proximally green or deep purple, distally red, crimson, scarlet, pink, pinkish purple, pinkish red, or yellow, sometimes rose magenta, red-orange, or orange, oblanceolate or obovate to oblong or lanceolate, (0–)3–5(–7)-lobed; lobes spreading to ascending, linear to obovate, long, proximal lobes at or arising below mid length, apices acute to obtuse, center lobe sometimes rounded. |
1.5–15 × 1.5–3 cm; bracts green throughout, or proximally green, distally white to rarely cream on apices, with a tuft of erect, white, soft hairs, especially when immature, narrowly lanceolate to narrowly ovate, 3–5-lobed; lobes ascending, linear to lanceolate, long, proximal lobes arising near base, apex obtuse to acute. |
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Pedicels | 0 mm or nearly so. |
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Corollas | straight or ± curved, 17–40 mm; tube 10–15 mm; beak long-exserted to subequal to calyx, adaxially green, 7–20 mm, puberulent, eglandular; abaxial lip deep green to reddish, brown, or deep purple, reduced, inconspicuous, included in calyx, 1.5–3 mm, 15–25% as long as beak; teeth ascending, green, 0.5–2 mm. |
straight or slightly curved distally, 12–25 mm; tube 18 mm; abaxial lip and beak exserted; beak adaxially white or pale yellow, 3.5–9 mm, hairs dense, spreading, medium length, obscuring surface; abaxial lip yellow, inflated, abruptly expanded, obpyramidal, pouches 3, central pouch slightly 2-lobed, 4–5 mm deep, 3–8 mm, 75–90% as long as beak; teeth erect, whitish to pale yellow, 1–2 mm. |
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Calyces | proximally pale or green, distally as in distal portion of bracts, 14–35 mm; abaxial and adaxial clefts 6–22 mm, 33–50% of calyx length, deeper than laterals, lateral 2–7 mm, 15–25% of calyx length; lobes lanceolate or oblong, apex acute to obtuse or rounded, sometimes curved upward. |
light green, lobes deep green, sometimes purple, 5.5–12 mm; abaxial and adaxial clefts 2.5–6.5 mm, 30–50% of calyx length, lateral 2–4.5 mm, 30–40% of calyx length; lobes linear to narrowly lanceolate or triangular, apex acuminate or acute. |
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2n | = 48, 72, 96. |
= 24. |
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Castilleja affinis |
Castilleja lasiorhyncha |
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Phenology | Flowering Jun–Jul. | |||||||||
Habitat | Springs, moist or wet meadows, flats, open forests. | |||||||||
Elevation | 1000–2500 m. (3300–8200 ft.) | |||||||||
Distribution |
CA; nw Mexico
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CA
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Discussion | Varieties 3 (3 in the flora). Castilleja affinis is highly variable and one of the more common paintbrushes at lower elevations in California, west of the Sierra Nevada, from the northern coast south to northern Baja California. Some recent authors (for example, M. Wetherwax et al. 2012) include C. litoralis of the Pacific Northwest coast as a subspecies of C. affinis, but due to the high polyploid nature of C. litoralis and its significantly closer morphological resemblance to C. miniata var. dixonii, that treatment is not followed here. Also see the comments under 62. C. litoralis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most populations of Castilleja lasiorhyncha are in the San Bernardino Mountains, with a few records in the adjacent Peninsular Ranges immediately to the south. The distal tufts of soft, pale hairs on the immature bracts are apparently unique in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 582. | FNA vol. 17, p. 618. | ||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Orthocarpus lasiorhynchus | |||||||||
Name authority | Hooker & Arnott: Bot. Beechey Voy., 154. (1833) | (A. Gray) T. I. Chuang & Heckard: Syst. Bot. 16: 657. (1991) | ||||||||
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