Castilleja affinis |
Castilleja chrysantha |
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California paintbrush, coast Indian paintbrush, coast paintbrush, Indian paintbrush |
Wallowa paintbrush, yellow paintbrush, yellow Wallowa Indian paintbrush, yellowish or common Wallowa paintbrush |
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Habit | Herbs, perennial, 1.4–6 dm; caudex woody; with a taproot. | Herbs, perennial, (0.5–)1–2(–5) dm; from a woody caudex; with a taproot. | ||||||||
Stems | few to many, erect to ascending, unbranched or branched at proximal nodes, sometimes with short, leafy axillary branches, hairs sparse to dense, spreading, short and long, soft to stiff, unbranched, sometimes branched, eglandular, sometimes stipitate-glandular. |
few to many, erect or ascending, often decumbent at base, unbranched, sometimes branched, hairs spreading, long, soft, mixed with dense, shorter stipitate-glandular ones. |
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Leaves | green or purplish, sometimes red-brown, linear or linear-oblong to broadly lanceolate, 2–13 cm, not or ± fleshy, margins wavy or plane, flat or involute, 0–5-lobed, sometimes with small secondary lobes, short, leafy axillary shoots common, usually conspicuous, apex acuminate or acute to rounded; lobes spreading, linear to lanceolate, lateral lobes almost as wide as mid blade, apex acute to rounded. |
green or purple-tinged, sometimes deep purple, broadly lanceolate, sometimes linear-lanceolate or narrowly oblong, 1.5–4.8 cm, not fleshy, margins plane, flat or involute, 0–3-lobed, apex acute; lobes ascending or spreading, linear to narrowly linear or narrowly lanceolate, short, apex acuminate. |
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Inflorescences | 3–25(–30) × 1.5–5 cm; bracts proximally green or deep purple, distally red, crimson, scarlet, pink, pinkish purple, pinkish red, or yellow, sometimes rose magenta, red-orange, or orange, oblanceolate or obovate to oblong or lanceolate, (0–)3–5(–7)-lobed; lobes spreading to ascending, linear to obovate, long, proximal lobes at or arising below mid length, apices acute to obtuse, center lobe sometimes rounded. |
3–17 × 1–3 cm; bracts greenish or pale yellow-green throughout, or proximally greenish or pale yellow-green, distally pale yellow to whitish, sometimes pink-purple, or pale, dull purplish, sometimes aging pink or yellow, often infused with light purple, rarely pink, ovate to broadly acute, (0–)3-lobed; lobes ascending, linear-lanceolate, oblong, or triangular, short, arising below mid length, apex acute to obtuse. |
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Pedicels | 0 mm or nearly so. |
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Corollas | straight or ± curved, 17–40 mm; tube 10–15 mm; beak long-exserted to subequal to calyx, adaxially green, 7–20 mm, puberulent, eglandular; abaxial lip deep green to reddish, brown, or deep purple, reduced, inconspicuous, included in calyx, 1.5–3 mm, 15–25% as long as beak; teeth ascending, green, 0.5–2 mm. |
straight, (16–)20–25 mm; tube 15–18 mm; beak slightly exserted, adaxially green to yellow, 5.5–8.5 mm; abaxial lip white, green and white, pink, or purple, ± prominent, appressed (proximally scarcely or not pouched), 3–5 mm, 67% as long as beak; teeth erect, white or pink with some purple or red, 1.5–2.5 mm. |
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Calyces | proximally pale or green, distally as in distal portion of bracts, 14–35 mm; abaxial and adaxial clefts 6–22 mm, 33–50% of calyx length, deeper than laterals, lateral 2–7 mm, 15–25% of calyx length; lobes lanceolate or oblong, apex acute to obtuse or rounded, sometimes curved upward. |
proximally green, pale with green veins, purple-tinged green, or purple, distally pale yellow, white, or purplish, 12–20 mm; abaxial and adaxial clefts 8–12 mm, 50% of calyx length, deeper than laterals, lateral 0.5–3 mm, 5–15% of calyx length; lobes short-triangular, apex obtuse, rounded, or truncate. |
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2n | = 48, 72, 96. |
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Castilleja affinis |
Castilleja chrysantha |
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Phenology | Flowering May–Oct. | |||||||||
Habitat | Flat, mesic meadows, dry talus and ridges, montane to alpine. | |||||||||
Elevation | 1100–2800 m. (3600–9200 ft.) | |||||||||
Distribution |
CA; nw Mexico
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OR
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Discussion | Varieties 3 (3 in the flora). Castilleja affinis is highly variable and one of the more common paintbrushes at lower elevations in California, west of the Sierra Nevada, from the northern coast south to northern Baja California. Some recent authors (for example, M. Wetherwax et al. 2012) include C. litoralis of the Pacific Northwest coast as a subspecies of C. affinis, but due to the high polyploid nature of C. litoralis and its significantly closer morphological resemblance to C. miniata var. dixonii, that treatment is not followed here. Also see the comments under 62. C. litoralis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja chrysantha is endemic to the mountains of northeastern Oregon, and its patterns of variation need further study. Most plants in the Blue Mountains are taller, more erect, and tolerate lower elevation habitats than populations around the type locality in the Wallowa Mountains. Plants with purplish inflorescences and longer hairs were described as C. ownbeyana and tend to favor drier talus and ridges than plants like the type, found in flat, mesic, montane to subalpine meadows. Hybrids between C. chrysantha and C. fraterna occur in Wallowa County. In the Wallowa Mountains, a recurrent and variable hybrid form between C. chrysantha and C. rhexiifolia was described as C. wallowensis Pennell. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 582. | FNA vol. 17, p. 596. | ||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | C. ownbeyana | |||||||||
Name authority | Hooker & Arnott: Bot. Beechey Voy., 154. (1833) | Greenman: Bot. Gaz. 48: 146. (1909) | ||||||||
Web links |