Castilleja affinis |
Castilleja ambigua |
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California paintbrush, coast Indian paintbrush, coast paintbrush, Indian paintbrush |
estuarine paintbrush, johnny-nip, owl-clover paintbrush, paint-brush owl-clover, paintbrush, paintbrush owl's-clover |
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Habit | Herbs, perennial, 1.4–6 dm; caudex woody; with a taproot. | Herbs, annual, sometimes biennial, 0.6–3.5 dm; with fibrous roots. | ||||||||||||||||||||
Stems | few to many, erect to ascending, unbranched or branched at proximal nodes, sometimes with short, leafy axillary branches, hairs sparse to dense, spreading, short and long, soft to stiff, unbranched, sometimes branched, eglandular, sometimes stipitate-glandular. |
few to many, sometimes solitary (var. meadii), ascending or erect, often decumbent proximally, unbranched or branched above base, hairs sparse to moderately dense, spreading, long, soft, eglandular, mixed with short stipitate-glandular hairs. |
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Leaves | green or purplish, sometimes red-brown, linear or linear-oblong to broadly lanceolate, 2–13 cm, not or ± fleshy, margins wavy or plane, flat or involute, 0–5-lobed, sometimes with small secondary lobes, short, leafy axillary shoots common, usually conspicuous, apex acuminate or acute to rounded; lobes spreading, linear to lanceolate, lateral lobes almost as wide as mid blade, apex acute to rounded. |
green or brownish in upland forms, linear-lanceolate to widely lanceolate, linear, elliptic, obovate, or oblong, rarely ovate or cup-shaped, (0.6–)0.8–5 cm, fleshy or not, margins plane, flat, 3–5(–7)-lobed, apex rounded to obtuse or acuminate, abaxial surface often stipitate-glandular, adaxial sometimes shiny, glabrous; lobes ascending to erect, linear to lanceolate, apex acute or obtuse. |
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Inflorescences | 3–25(–30) × 1.5–5 cm; bracts proximally green or deep purple, distally red, crimson, scarlet, pink, pinkish purple, pinkish red, or yellow, sometimes rose magenta, red-orange, or orange, oblanceolate or obovate to oblong or lanceolate, (0–)3–5(–7)-lobed; lobes spreading to ascending, linear to obovate, long, proximal lobes at or arising below mid length, apices acute to obtuse, center lobe sometimes rounded. |
1.5–9(–13) × 1–4 cm; bracts proximally green, rarely brownish purple, distally white, cream, pink, or purple on apices, lanceolate, oblong, or ovate, (0–)3–5(–7)-lobed; lobes ascending or divaricate-ascending, oblong to linear, short to long, arising below or above mid length, central lobe apex usually rounded to truncate, others acute to obtuse. |
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Pedicels | 0 mm or nearly so. |
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Corollas | straight or ± curved, 17–40 mm; tube 10–15 mm; beak long-exserted to subequal to calyx, adaxially green, 7–20 mm, puberulent, eglandular; abaxial lip deep green to reddish, brown, or deep purple, reduced, inconspicuous, included in calyx, 1.5–3 mm, 15–25% as long as beak; teeth ascending, green, 0.5–2 mm. |
straight, 14–24 mm; tube 11–21 mm, expanded distally; abaxial lip and beak usually exserted, beak straight or slightly curved, adaxially white, yellow, or pink, sometimes green or purplish, (1–)4–7 mm, sparsely to densely short-hairy; abaxial lip pale to bright yellow, sometimes becoming pink, orange, or red after anthesis, with red-brown or purple spots at base of each tooth and sometimes at base of each pouch, conspicuous, pouches 3, ± prominent, divergent, saccate, 2–7 × 3–7 mm, (33–)60–75(–90)% as long as beak; teeth erect, sometimes spreading, white, pink, purple, or green, often with whitish bases, 1–3 mm. |
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Calyces | proximally pale or green, distally as in distal portion of bracts, 14–35 mm; abaxial and adaxial clefts 6–22 mm, 33–50% of calyx length, deeper than laterals, lateral 2–7 mm, 15–25% of calyx length; lobes lanceolate or oblong, apex acute to obtuse or rounded, sometimes curved upward. |
green to pale yellowish green or tipped with white or purple, 8–23 mm; abaxial and adaxial clefts 5–9 mm, 33–50(–67)% of calyx length, lateral 2–5.5 mm, 20–40% of calyx length; lobes linear or narrowly lanceolate to oblong or ± triangular, apex acuminate or narrowly acute to obtuse or rounded. |
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Filaments | glabrous. |
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2n | = 48, 72, 96. |
= 24. |
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Castilleja affinis |
Castilleja ambigua |
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Distribution |
CA; nw Mexico
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CA; OR; WA; BC
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Discussion | Varieties 3 (3 in the flora). Castilleja affinis is highly variable and one of the more common paintbrushes at lower elevations in California, west of the Sierra Nevada, from the northern coast south to northern Baja California. Some recent authors (for example, M. Wetherwax et al. 2012) include C. litoralis of the Pacific Northwest coast as a subspecies of C. affinis, but due to the high polyploid nature of C. litoralis and its significantly closer morphological resemblance to C. miniata var. dixonii, that treatment is not followed here. Also see the comments under 62. C. litoralis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Castilleja ambigua is a complex species, treated here with four varieties, though many localized variations exist among populations, and these are still incompletely understood. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 582. | FNA vol. 17, p. 584. | ||||||||||||||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||||||||||||||
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Name authority | Hooker & Arnott: Bot. Beechey Voy., 154. (1833) | Hooker & Arnott: Bot. Beechey Voy., 154. (1833) | ||||||||||||||||||||
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