Cascadia nuttallii |
Cascadia |
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Nuttall's saxifrage |
saxifrage |
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Habit | Herbs, annual; herbage glabrous or sparsely red stipitate-glandular. | |
Stems | branched proximal to middle, slender. |
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Flowering stems | trailing or suberect, leafy, 4–40 cm. |
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Leaves | largest along middle of stems; petiole distinct, 1–6 mm; blade 3–20 mm. |
cauline; stipules absent; petiole glabrous; blade ovate to lanceolate, unlobed, base cuneate or attenuate to rounded, margins 3-toothed apically or not, ultimate margins entire, (without lime-secreting hydathodes), apex acute; venation palmate or pinnate. |
Inflorescences | poorly defined, usually paniculiform or racemiform thyrses, sometimes solitary flowers, 2–4-flowered, bracteate. |
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Pedicels | filiform. |
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Flowers | sepals erect, triangular, apex acute; petals elliptic to obovate, 3–6 mm, much longer than sepals. |
hypanthium adnate to proximal 1/3 of ovary, free from ovary 0 mm, adnate portion increasing along with attached ovary at maturity to constitute 2/3–4/5 of fruit, green, (0.8–2 mm); sepals 5, green; petals 5, white; nectary disc present; stamens 10, (distinct); filaments flattened, slightly widened at base; ovary semi-inferior, 2-locular, carpels connate in proximal 1/3 or less; placentation axile, appearing marginal above point of connation of ovaries; styles 2; stigmas 2. |
Capsules | 2–5 mm. |
folliclelike, 2-beaked. |
Seeds | brown, oblong, spiny in longitudinal rows. |
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x | = 8. |
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2n | = 16. |
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Cascadia nuttallii |
Cascadia |
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Phenology | Flowering spring–summer. | |
Habitat | Wet, shaded cliffs and ledges, sometimes coastal or near waterfalls, mossy mats, wet rocks, moist crevices | |
Elevation | 0-1200 m (0-3900 ft) | |
Distribution |
CA; OR; WA
|
w United States |
Discussion | Cascadia nuttallii is found from the coastal mountains to the western slopes of the Cascade Range, from extreme northwestern California to southwestern Washington. The accepted species name was validated in a correction slip attached to reprints of Johnson’s article describing Cascadia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 1: w United States. Johnson placed Saxifraga nuttallii in his monotypic genus Cascadia based on the unusual habit, free carpels, and spiny seeds. Molecular phylogenetic data (M. E. Mort and D. E. Soltis 1999; Soltis et al. 2001) placed Cascadia as sister to the southern South American (Tierra del Fuego) Saxifragodes D. M. Moore, both sister to Micranthes. Mort and Soltis considered the ovary of Cascadia to be superior because the two carpels are distinct to their bases; the hypanthium, fused to each carpel, gives the ovaries a semi-inferior appearance. Ovules in Cascadia are bitegmic, as in Saxifraga; those of Micranthes are usually unitegmic. Species 1 (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 8, p. 49. | FNA vol. 8, p. 48. |
Parent taxa | Saxifragaceae > Cascadia | Saxifragaceae |
Subordinate taxa | ||
Synonyms | Saxifraga nuttallii | |
Name authority | (Small) A. M. Johnson: Corrig. 1927 , | A. M. Johnson: Amer. J. Bot. 14: 38, figs. 1, 2. 1927 , |
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