Caryophyllaceae subfam. Polycarpoideae |
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| Habit | Herbs [small shrubs], winter annual, annual, or perennial; taprooted, not rhizomatous. |
| Stems | prostrate to erect, simple or branched. |
| Leaves | opposite or sometimes appearing whorled, bases connate or not, sometimes petiolate or often sessile, stipulate; stipules ovate or deltate to lanceolate or bristlelike, scarious; blade subulate or subtriangular to linear and threadlike or spatulate to ovate or orbiculate, seldom succulent. |
| Inflorescences | terminal or axillary cymes, or flowers solitary; bracts scarious or absent; involucel bracteoles absent. |
| Pedicels | present or flowers sessile. |
| Flowers | bisexual or rarely unisexual; perianth and androecium hypogynous or perigynous; hypanthium absent or dish- or cup-shaped; sepals 5, distinct or sometimes connate proximally, hooded (Drymaria, Polycarpon) or not, awned (Polycarpon) or not; petals absent or (3–)5, blade clawed (Drymaria) or not, auricles absent, coronal appendages absent, blade apex entire, erose, or 2(–4)-fid, sometimes emarginate; stamens (1–)3–5(–10), in 1 or 2 whorls, usually arising from base of ovary or from rim of hypanthium (Spergula, Spergularia); staminodes absent; ovary 1-locular; styles 1 or 3, occasionally 2 or 5, distinct or sometimes connate proximally; stigmas 3, occasionally 2 or 5. |
| Fruits | capsules, opening by 3 or 5, occasionally 4 valves; carpophore sometimes present. |
| Seeds | 3–150+, whitish or tan to often brown or black, ± triangular, pyriform, or reniform to circular, subglobose or laterally compressed to angular (Polycarpon); embryo peripheral, curved or rarely annular to spirally curved (Spergula). |
| x | = [7], 8, 9, (11), 12. |
Caryophyllaceae subfam. Polycarpoideae |
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| Distribution | w North America (including Mexico); w South America; Europe (Mediterranean region); Asia (Mediterranean region); Africa (Mediterranean and tropical regions); less diverse in temperate areas |
| Discussion | Genera 16, species ca. 210 (7 genera, 28 species in the flora). More commonly recognized as a tribe, Polycarpoideae is characterized by the presence of stipules, well-developed petals, and capsules. It is a relatively small group, with about four-fifths of the species in Drymaria, Polycarpaea, and Spergularia. Although clearly stipulate, Spergularia clusters with members of Alsinoideae in recent molecular studies (M. Nepokroeff et al. 2002; R. D. Smissen et al. 2002). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| Name authority | Tanfani: in F. Parlatore, Fl. Ital. 9: 623. (1892) |
| Source | FNA vol. 5, p. 9. |
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