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limestone forest sedge, Willdenow's sedge

arctic sedge, capitate sedge, carex capité

Culms

erect, (4.7–)7.1–14.4 cm, 1/5–2/5 of total plant height.

Leaves

basal sheaths pale brown to dark brown;

blades green, much exceeding culms, 1.2–4.6 mm wide, herbaceous, margins green, smooth or scabrous.

Perigynia

(3–)4–9 per spike, pale green to pale brown, often speckled reddish brown, narrowly ovoid or lance-ovoid, (7–)7.5–8.9(–10.8) × 1.5–2.3 mm, tightly enveloping achenes, apex gradually tapering;

beak (3.6–)4.1–5(–6.4) mm, scabrous.

2–4 × 1.5–1.8 mm, distal margins smooth;

beak smooth or sparingly serrulate.

Achenes

brown, oblong, (2.1–) 2.5–3.1(–3.3) × (1.4–)1.5–1.9(–2.1) mm.

Lateral

spikes 0–2(–3), basal, on erect or spreading (rarely drooping) peduncles.

Terminal

spikes with staminate portion 7–56-flowered, (5.4–) 9.6–20.1 × 0.7–1.6 mm; pistillate portion spike 2–4(–6)-flowered.

Pistillate

scales green, margins hyaline, white, tinged reddish brown, (0.9–)1.2–2.4 mm wide, not more than 1.5 times as wide and not concealing perigynia, apex attenuate into green awns;

distal scales with hyaline margins 0.3–0.8 mm wide, apex acuminate, awnless, or scabrous-awned to 5.7 mm.

scales ovate, as long as or much shorter than the perigynia, margins narrowly or broadly hyaline.

Staminate

scales green or tinged with pale brown, ovate, 2.5–4.1 × 1.6–2 mm, margins free but enfolding scales above, hyaline, white, apex obtuse, erose.

Anthers

1.1–1.4 mm.

Stigmas

filiform, flexuous, long, thin, strongly papillose.

2n

= 50.

Carex superata

Carex capitata

Phenology Fruiting spring–summer (late Mar–mid Jul). Fruiting Jul–Aug.
Habitat Acidic to neutral clays, often high in potassium and magnesium, in moist to dry mesic, open, deciduous forests, ravines Mires and heaths of the boreal forest with disjunct occurrences southward in the alpine zone of eastern and western mountain ranges, primarily on calcareous substrates
Elevation 50–500 m (200–1600 ft) 20–3400 m (100–11200 ft)
Distribution
from FNA
AL; FL; GA; KY; MS; SC; TN; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; CO; MT; NH; NV; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Mexico; South America; Greenland; Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

A. Cronquist (1969) placed Carex capitata in Utah; the species is not listed by B. J. Albee et al. (1988), nor have specimens been seen from there.

In Scandinavia (L. Reinhammar 1999) and Russia (T. V. Egorova 1999), Carex arctogena is distinguished from C. capitata on ecologic and morphologic grounds and also by allozymes at the rank of species. In North America the distinctions are not clear; two taxa can be observed, but there are also numerous specimens of uncertain determination. When making new collections, it is important to evaluate the differences seen elsewhere and to pay close attention to habitat and habit (see below). Separate status at some rank may be appropriate for the taxon “arctogena” in North America too. If C. antarctogena Roivainen from Tierra del Fuego is placed within C. capitata as D. M. Moore and A. O. Chater (1971) and D. M. Moore (1983) have done, then the species occurs at both the northern and the southern extremes of the Americas.

Carex capitata and C. arctogena differ in habitat (boreal mires versus alpine heaths), habit (mat-forming versus tufted), and morphology of the pistillate scales (much shorter and narrower than perigynia and with narrow hyaline margins versus as long as perigynia and with broad hyaline margins) and perigynia (beak gradually formed and smooth versus beak and may be sparingly serrulate).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 560. FNA vol. 23, p. 569.
Parent taxa Cyperaceae > Carex > sect. Phyllostachyae Cyperaceae > Carex > sect. Capituligerae
Sibling taxa
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. aquatilis, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. calcifugens, C. californica, C. canescens, C. capillaris, C. capitata, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chalciolepis, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. echinata, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. gracillima, C. granularis, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lupulina, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vaginata, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. aquatilis, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. calcifugens, C. californica, C. canescens, C. capillaris, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chalciolepis, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. echinata, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. gracillima, C. granularis, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lupulina, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. superata, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vaginata, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica
Synonyms C. willdenowii var. megarrhyncha C. arctogena, C. capitata var. arctogena, C. capitata subsp. arctogena
Name authority Naczi: Amer. J. Bot. 85: 443, figs. 1, 2, 4, 5, 6(left). (1998) Linnaeus: Syst. Nat. ed. 10, 2: 1261. (1759)
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