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carex houblon, hop sedge

Habit Plants loosely cespitose or not, short-rhizomatous. Plants cespitose or not, rarely colonial, short to long rhizomatous.
Culms

solitary or not, erect, 20–100(–130) cm.

purplish or reddish, rarely brown at base.

Leaves

4–8;

basal sheaths reddish to brownish;

sheath of distal leaf (1.3–)1.7–10(–25) cm;

ligules triangular, 3.5–18 mm;

blades 15–64 cm × 4–15 mm.

basal sheaths not fibrous;

sheath fronts membranous;

sheaths and larger leaves distinctly septate-nodulose;

blades V-shaped in cross section when young, glabrous.

Inflorescences

4–40 cm;

peduncles of proximal spikes 0.5–20 cm, basal 2 peduncles 1–20 cm apart of terminal spike 0.5–6(–7) cm, shorter than or exceeding distal pistillate spike by 2 cm or less;

bracts leafy, sheaths 0.5–15 cm, blades 13–55 cm × 3–11 mm.

racemose, with 2–6(–9) spikes;

proximal bracts leaflike, sheathless or sheath less than 4 mm, shorter or longer than diam. of stem;

lateral spikes pistillate or the distal 1(–5) staminate or androgynous, globose, ovoid, or cylindric, pedunculate, prophyllate;

terminal spike staminate.

Spikes

proximal pistillate spikes (1–)2–5, distal spikes usually ± crowded, ascending, densely (4–)8–80-flowered, ovoid to cylindric, 1.5–6.5 × 1.3–3 cm;

terminal staminate spikes 1(–2), 1.5–8.5 cm × 1–5 mm.

Perigynia

ascending, strongly 13–22-veined, ± stipitate, lance-ovoid, 11–19 × 3–6 mm, shiny, glabrous;

beak conic, 6–10 mm.

ascending to spreading, distinctly 15–20-veined, sometimes stipitate, inflated, ovate, ± round in cross section, 10–20 mm, base cuneate to rounded, apex tapering or abruptly beaked, glabrous or sparsely hairy;

beak conspicuously bidentate, teeth not more than 1 mm.

Achenes

stipitate, rhomboid, flat to concave faces, angles smoothly thickened, not pointed or nobbed, 3–4(–4.5) × 1.7–2.6(–2.8) mm;

style same texture as achene.

trigonous, smaller than bodies of perigynia;

style persistent.

Pistillate

scales 1–7-veined, lanceolate to lanceolate-ovate, 6–15 × 1–2.7 mm, apex acute to awned, awns rough, to 6 mm.

Anthers

3, 3–5.5 mm.

Proximal

pistillate scales with apex obtuse to acuminate, often awned.

Stigmas

3.

Carex lupulina

Carex sect. Lupulinae

Phenology Fruiting late spring–summer.
Habitat Wet, deciduous or mixed swamp forests, forest openings, wet meadows
Elevation 0–700 m (0–2300 ft)
Distribution
from FNA
AL; AR; CT; DC; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
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[BONAP county map]
e North America
Discussion

Occasional specimens have been seen that appear to be sterile hybrids of Carex lupulina with C. retrorsa or C. lurida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 6 (6 in the flora).

Members of Carex sect. Lupulinae have the largest perygynia of all Carex. Mature achenes are critical for identification of some species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Sheath of the distal nonbracteal leaf 0–1.5(–2.5) cm; beak of perigynium 1.5–4.2 mm; achenes elliptic or obovate; spikes globose to short-ovoid.
→ 2
1. Sheath of the distal nonbracteal leaf usually 1.7 cm or longer; beak of perigynium 4.5–10 mm; achenes rhombic or nearly triangular; spikes ovoid to cylindric.
→ 3
2. Perigynia radiating out in all directions to form globular spike, rhombic-ovoid, base cuneate, 8–35 per spike.
C. grayi
2. Perigynia ascending to spreading or, sometimes, the basalmost reflexed to form an ovoid to obovoid spike, lanceoloid to ovoid, base convex, 1–12(–20) per spike.
C. intumescens
3. Achenes distinctly wider than long, widest beyond midle; perigynia stiffly spreading at right angles to rachis.
C. gigantea
3. Achenes as wide as long as or longer, widest near middle; perigynia ascending.
→ 4
4. Angles of achenes pointed, often knobbed, with hard, nipplelike points; achenes (2.2–)2.4–3.4 mm wide, often nearly as wide as long.
C. lupuliformis
4. Angles of achenes smoothly curved, not pointed or knobbed; achenes 1.7–2.6(–2.8) mm wide, longer than wide.
→ 5
5. Staminate peduncle 0.5–6(–7) cm, shorter than to exceeding distal pistillate spike by no more than 2 cm; plants loosely cespitose or not, short-rhizomatous.
C. lupulina
5. Staminate peduncle (3–)6–18 cm, usually exceeding distal pistillate spike by 2–12 cm; plants loosely colonial, long-rhizomatous.
C. louisianica
Source FNA vol. 23. FNA vol. 23. Author: A. A. Reznicek.
Parent taxa Cyperaceae > Carex > sect. Lupulinae Cyperaceae > Carex
Sibling taxa
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. aquatilis, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. calcifugens, C. californica, C. canescens, C. capillaris, C. capitata, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chalciolepis, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. echinata, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. gracillima, C. granularis, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. superata, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vaginata, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica
Subordinate taxa
C. gigantea, C. grayi, C. intumescens, C. louisianica, C. lupuliformis, C. lupulina
Name authority Willdenow: Sp. Pl. 4(1): 266. (1805) Tuckerman ex J. Carey: Carices North. U.S., 562. (1847)
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