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carex granuleux, limestone-meadow sedge, meadow sedge
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calcium-hating sedge
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Plants densely cespitose; rhizome internodes 0.2–7 mm, 1.2–2.2 mm thick. |
short or inconspicuous. |
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in dense tufts, 13–70(–100) cm. |
sparingly purple-red to 2.7–5.2(–5.6) cm high at base, 14–37 cm; longest vegetative shoots 23–45 cm, (0.8–)1.4–1.6 times as tall as culms. |
not green, usually glaucous, cauline blades 2–27 cm × 1.2–5.3 mm. |
nonbasal sheaths glabrous; blades green, widest blades 2.6–4 mm wide, smooth abaxially. |
|
19–34 cm, (0.88–)0.94–0.99 of culm height; internode between proximalmost scales in proximalmost spike 3–4.3 mm; internode between distal lateral spikes (1–)2.6–12 cm; longest internode (per plant) between distal lateral spikes 5.2–12 cm; peduncle of terminal spike 3.1–54(–73) mm, longest per plant 17–54(–73) mm; proximal bract sheaths tight, apex concave or truncate; ligules 1–4.9(–9.6) mm; blades uniformly green, blade of distalmost lateral bract exceeding terminal spike, longest (per plant) bract blade of distalmost lateral spike 6.1–15 cm. |
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4–5 spikes, widely separate or distal 2–3 spikes overlapping, proximalmost spike usually basal; lateral spikes pistillate, with 2–8 perigynia, longest spikes per plant with 5–8, 11–17 × 4.5–8.5 mm, ratio of longest lateral spike length (in mm) to flower number = 2.2–2.4; terminal spike 8–35 × 1.4–2.5 mm, longest per plant 14–35 mm, exceeding distlmost lateral spike. |
olive to yellowish green or at age yellowish brown, elliptic- or oblong-ovoid to obovoid, 2.2–3.1(–3.7) × 1.2–1.8(–2) mm, 1.4–2.2(–2.4) times as long as thick; beak 0.1–0.3 mm, orifice nearly entire. |
distichously imbricate, prominently 48–63-veined, obovate or narrowly obovate, obtusely trigonous in cross section, (3.7–) 3.9–5 × 1.6–2(–2.1) mm, 2.1–2.5(–2.7) times as long as wide, 1.5–2 times as long as achene bodies, base gradually tapered, apex contracted; beak straight or excurved 10–30°, (0.4–)0.6–1 mm, 0.08–0.21 of perigynium length. |
1.8–2.3 × 1–1.4 mm. |
, 2.6–3.4 × 1.4–1.9 mm, body 2–2.6 mm, tightly enveloped by perigynia; stipe straight, 0.2–0.5(–0.7) mm; beak straight, (0.1)–0.2(–0.3) mm. |
peduncle of terminal spike 0.1–3.5 cm; bracts (middle and distal ones) overtopping spikes; ligule of proximal bract (2–)3–17.5(–26) mm; longest bract blade (per plant) of distal lateral spike 4.1–15.8 cm. |
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spikes usually arising from distal 1/2 of culms, 6–27(–32) × 3–6 mm. |
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spike 6–35(–43) mm, shorter than to barely exceeding and overlapping distal lateral spike. |
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scales ovate or ovateoblong, 1.4–2.5(–2.9) × 0.8–1.4 mm. |
scales 3.6–5.7 × 1.8–2.1(–2.4) mm, body (2.2–)2.5–3 mm, midrib prolonged as awn 01–3.2 mm. |
scales with apex acute to acuminate or cuspidate. |
scales 4.3–5.8 × 1.4–1.9 mm, apex usually acute, occasionally obtuse, awnless. |
1.6–2.9 mm. |
3.8–4.2 mm. |
= 36, 38, 40, 42. |
|
|
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|
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| Fruiting late spring–early summer (late Apr–Jul). |
Fruiting spring. |
| Low, open ground, meadows, fens, glades, or shores, ditches, moist woods, and bottomland swamps, especially along borders, clearings, streams, trails, usually in clayey or marly soils, frequently weedy in limestone districts |
Mesic deciduous forests, in sandy loams and sands, usually on stream banks slopes |
| 0–700 m (0–2300 ft) |
0–40 m (0–100 ft) |
|
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; PA; SD; TN; TX; VA; VT; WI; WV; WY; MB; NB; NS; ON; QC; SK
|
FL; GA; NC; SC |
Carex granularis is widely distributed and common across the eastern half of the United States and southeastern Canada, but not on the Coastal Plain or in the high Appalachians. It varies not only in the dimensions and ultimate overall shape of the perigynia but also in less tangible features, such as texture, color, and amount of red speckling on the perigynia, prominence of perigynia veins, and papillosity of achene surface. One phase distinguished by the smaller, slightly less inflated, more ellipsoid perigynia with more pointed apices and finer veins has often been treated as C. haleana or C. shriveri; it intergrades rather freely with typical C. granularis. Because of the subtlety of the distinctions, the intergradation in morphology, and the difficulties presented by specimens in immature stages or distorted during preparation, we maintain C. granularis as a single variable species without infraspecific taxa. An apparent hybrid between C. granularis and C. flaccosperma Dewey or C. glaucodea Tuckerman ex Olney is known from one locality in Arkansas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Carex calcifugens is rare, local, and usually occurs in small populations. It frequently associates with C. abscondita and C. striatula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 23, p. 441. |
FNA vol. 23, p. 453. |
| Cyperaceae > Carex > sect. Granulares |
Cyperaceae > Carex > sect. Griseae |
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. aquatilis, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. calcifugens, C. californica, C. canescens, C. capillaris, C. capitata, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chalciolepis, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. echinata, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. gracillima, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lupulina, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. superata, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vaginata, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica |
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. aquatilis, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. californica, C. canescens, C. capillaris, C. capitata, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chalciolepis, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. echinata, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. gracillima, C. granularis, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lupulina, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. superata, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vaginata, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica |
| C. granularis var. haleana, C. haleana, C. rectior, C. shriveri |
|
| Muhlenberg ex Willdenow: Sp. Pl. 4(1): 279. (1805) |
Naczi: Novon 12: 512, fig. 2. (2002) |
| |
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