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carex filiforme, graceful sedge

carex engaîné, sheath sedge

Habit Plants densely cespitose. Plants colonial, long-rhizomatous.
Culms

dark maroon at base; flowering stems 0.5–1.1 mm thick, 20–90 cm, much longer than leaves at maturity, glabrous, but often scabrous within inflorescence.

7–60 cm, ± smooth distally.

Leaves

sheaths glabrous or short-pubescent, basal few bladeless and maroon, others grading from maroon to green on back, pale brown-hyaline, red dotted and sometimes pubescent near apex;

blades flat, 3–9 mm wide, glabrous, minutely scabrous on margins distally.

proximal sheaths bladeless, pale brown, slender, 0.7–2.8 mm diam., not or only very sparsely papillose;

ligules 2–4.5 mm, longer than wide;

blades of vegetative culms 5–40 cm × 1.5–4 mm;

those of fertile culms green, flat, less than 10 cm.

Inflorescences

peduncles of lateral spikes slender, to 40 mm, equaling spikes or usually somewhat shorter, scabrous;

peduncle of terminal spike to 13 mm, glabrous or scabrous;

proximal bracts shorter than inflorescences;

sheaths 20–25 mm;

blades 3–4 mm wide.

6.5–25 cm, 2–5 times longer than proximal bracts;

proximal bracts 2.5–8 cm, sheath 1–3.5 cm, blade 1–1.5 cm;

pistillate spikes linear, 0.8–3 cm × 3–6.5 mm.

Perigynia

green, usually red dotted, 2-ribbed and finely 8–12-veined, veins more obvious near base, ellipsoid to oblong-ovoid, loosely enveloping achenes, 2–3.7 × 1.3–1.6 mm, membranous, acute at base, tapering gradually to acute beakless apex, glabrous.

3–15, the proximal loosely arranged, green to dark brown, often obscurely veined, 3.5–5 × 1.5–2.2 mm, smooth;

beak slender, cylindric, (0.4–)0.6–1.8(–2.2) mm.

Achenes

substipitate, 1.2–2.6 × 1–1.2 mm.

light to dark brown, 2.2–3 × 1.2–1.5 mm.

Lateral

spikes 2–5, 1 per node, well separated, drooping at maturity, pistillate with 10–45 perigynia attached 2 mm apart, linear, 10–70 × 2–3.5 mm.

Terminal

spike gynecandrous, 10–60 × 1–3 mm.

Pistillate

scales white-hyaline tinged with chestnut, broad green midrib often red dotted, sometimes scabrous, ovate to oblong, shorter than mature perigynia, apex rounded or cuspidate with ciliate awn less than 1 mm, otherwise glabrous.

scales purple tinged, apex broadly acute or obtuse.

2n

= 50, 52, 54.

= 32.

Carex gracillima

Carex vaginata

Phenology Fruiting late spring–mid summer. Fruiting early–mid summer.
Habitat Mesic to dry deciduous forests, including edges and openings, mixed conifer-hardwood forests, coniferous swamps, thickets, meadows, roadsides Calcareous swamps, boggy thickets and woods
Elevation 30–1600 m (100–5200 ft)
Distribution
from FNA
AL; AR; CT; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; ND; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; MB; NB; NL; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; ME; MI; MN; MT; NH; NY; VT; WI; AB; BC; MB; NB; NL; NT; NU; ON; QC; SK; YT; Greenland; Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

Two varieties of Carex gracillima have been named based on specimens with shorter stems, narrower leaves and smaller perigynia. Although detailed studies are lacking, these plants are more likely to be phenotypic responses to stressful conditions and do not warrant taxonomic recognition. Carex gracillima is common in a variety of forest habitats and has an intermediate blooming time among the eastern forest sedges. As a result, sterile hybrids with other species appear to be quite frequent and include: C. gracillima × aestivalis (= C. ×aestivaliformis), C. gracillima × hirtifolia (= C. ×sullivantii), C. gracillima × hirsutella, C. gracillima × virescens, C. gracillima × swanii, C. gracillima × castanea, and C. gracillima × arctata. Although none of the hybrids has been confirmed experimentally, at least the first four exhibit clear morphologic intermediacy and co-occur with the putative parental species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

M. L. Fernald (1950) pointed out that North American plants sometimes segregated as Carex saltuensis do not appear to be sufficiently distinct to warrant recognition as a separate species. T. V. Egorova (1999) recognized both C. saltuensis and C. vaginata, with two subspecies, in North America. More research is clearly needed on this complex circumpolar entity. The inclusion of C. vaginata in sect. Paniceae is uncertain. Unlike other members of the group, C. vaginata lacks papillae on abaxial leaf epidermis, peduncle, and perigynium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 428.
Parent taxa Cyperaceae > Carex > sect. Hymenochlaenae Cyperaceae > Carex > sect. Paniceae
Sibling taxa
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. aquatilis, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. calcifugens, C. californica, C. canescens, C. capillaris, C. capitata, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chalciolepis, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. echinata, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. granularis, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lupulina, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. superata, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vaginata, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. aquatilis, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. calcifugens, C. californica, C. canescens, C. capillaris, C. capitata, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chalciolepis, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. echinata, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. gracillima, C. granularis, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lupulina, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. superata, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica
Synonyms C. gracillima var. macerrima C. saltuensis
Name authority Schweinitz: Ann. Lyceum Nat. Hist. New York 1: 66. (1824) Tausch: Flora 4: 557. (1821)
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