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bristle fruit sedge, prickly sedge, star sedge

Holm sedge

Habit Plants densely cespitose.
Culms

10–90(–135) cm.

20–50 cm, smooth.

Leaves

3–6 per culm;

sheaths tight, inner band hyaline, 1.8–2.5 cm, apex concave, glabrous;

ligules 0.6–2.5(–4.5) mm, rounded to ± acute;

blades plicate, 5–40 cm × 0.7–3.3(–3.8) mm, widest leaf 1–3.3(–3.8) mm wide.

2.5–5 mm wide.

Inflorescences

0.7–7.8 cm;

spikes (2–)3–8;

lateral spikes pistillate often with few staminate flowers proximally, 3–15.5 mm, sessile, staminate portion 7-flowered, to 8.2 mm, pistillate portion 3–32-flowered, 2.5–11 mm;

basal 2 spikes 1.7–42 mm apart;

terminal spike gynecandrous, 5–24 mm, staminate portion 2–17-flowered, 2–16.5 × 0.9–2 mm, pistillate portion 4–26-flowered, 2.6–9 × 4.9–9.1 mm.

proximal bracts shorter than or exceeding inflorescences;

spikes contiguous, overlapping, distinct or the proximal separate and pendent, elongate, 10–25 × 6–10 mm;

proximal long-pendunculate;

lateral 3–6 spikes pistillate or, rarely, gynecandrous;

terminal spike gynecandrous.

Perigynia

spreading to reflexed, castaneous to dark brown, 2–14-veined abaxially, 0–12-veined adaxially over achene, lanceolate to ovate, (2.65–)2.9–4.75 × 0.8–2.1 mm, (1.7–)1.8–3.2(–3.6) times as long as wide;

beak (0.85–)0.95–2 mm, (0.4–)0.45–0.86 length of body, serrulate, teeth (0–)0.15–0.5 mm.

ascending, purple-brown with green or pale yellow margins, veinless, broadly ovate, 3–4 × 2.25–3.5 mm, apex abruptly beaked, distally papillose;

beak 0.3–0.5 mm, entire or shallowly bidentate, smooth.

Pistillate

scales ovate, (1.4–)2.1–3.1 × 0.7–2.3 mm, apex acute to sometimes obtuse.

scales chestnut or brownish black with hyaline margins, lanceolate, equaling or longer than and narrower than perigynia, midvein same color as body, inconspicuous, slightly raised, prominent.

Staminate

scales ovate-lanceolate to ovate, 1.3–3.8 × 0.8–1.7 mm, base clavate, apex acute.

Anthers

0.8–1.6(–2) mm.

Achene(s)

ovate-lanceolate to rhombic-ovate, 1.3–2.1 × 0.8–1.55 mm.

filling proximal 1/2 or less of perigynium.

2n

= 58.

Carex echinata

Carex chalciolepis

Phenology Fruiting Jun–Aug.
Habitat Subalpine and alpine meadows
Elevation 3000–4000 m (9800–13100 ft)
Distribution
from FNA
AK; CA; CO; CT; DC; DE; IA; ID; IL; IN; MA; MD; ME; MI; MN; MT; NC; ND; NH; NJ; NV; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SK; Mexico; Central America; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; MT; NM; NV; UT
[WildflowerSearch map]
Discussion

Subspecies 3 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Carex chalciolepis has pendent spikes with an almost shaggy appearance from pistillate scales typically longer than the perigynia. Similar forms are found among C. aterrima Hoppe in eastern Siberia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences lax to dense, usually 2.5–8 cm; distance between basal 2 spikes usually longer than basal spike.
subsp. echinata
1. Inflorescences very dense, 1.2–2.5(–4) cm; distance between basal 2 spikes usually less than length of basal spike.
→ 2
2. Perigynia 2.9–3.6(–4) mm, often veinless over achene on adaxial surface; widest leaves 1–2.4(–2.7) mm wide; widespread.
subsp. echinata
2. Perigynia (3.1)3.5–4.75 mm, 2–12-veined over achene on adaxial surface; widest leaves (1.7–)2.3–3.3(–3.8) mm wide; west coast.
subsp. phyllomanica
Source FNA vol. 23, p. 330. FNA vol. 23.
Parent taxa Cyperaceae > Carex > sect. Stellulatae Cyperaceae > Carex > sect. Racemosae
Sibling taxa
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. aquatilis, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. calcifugens, C. californica, C. canescens, C. capillaris, C. capitata, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chalciolepis, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. gracillima, C. granularis, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lupulina, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. superata, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vaginata, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. aquatilis, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. calcifugens, C. californica, C. canescens, C. capillaris, C. capitata, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. echinata, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. gracillima, C. granularis, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lupulina, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. superata, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vaginata, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica
Subordinate taxa
C. echinata subsp. echinata, C. echinata subsp. phyllomanica
Synonyms C. atrata var. chalciolepis, C. atrata var. larimerana, C. chalciolepis var. larimerana, C. heteroneura var. chalciolepis
Name authority Murray: Prodr. Stirp. Gott., 76. (1770) T. Holm: Amer. J. Sci. 166: 28, figs. 1–5. (1903)
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