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Photo is of parent taxon

calcareous water sedge, carex subétroit

water sedge

Habit Plants not cespitose.
Culms

35–100 cm, angles glabrous or scabrous.

20–150 cm.

Leaves

basal sheaths leafless, usually red-brown;

blades amphistomic, 5–8 mm wide, papillose on both surfaces.

basal sheaths red-brown or brown;

sheaths of proximal leaves glabrous, fronts with indistinct spots, veinless, apex U-shaped;

blades usually amphi-stomic, papillose on both surfaces.

Inflorescences

peduncle of proximal spike not more than 4 cm;

proximal bract longer than inflorescence, 4–8 mm wide.

proximal bract longer than inflorescence.

Spikes

erect;

proximal 2–6 spikes pistillate, 4–6 cm × 4–7 mm, base cuneate or attenuate;

terminal 1–3 spikes staminate.

erect;

proximal 2–6 spikes pistillate;

terminal (1–)2–4 spikes staminate.

Perigynia

pale brown, veinless, obovoid, 2.6–3.2 × 1.5–2.2 mm, apex rounded;

beak pale brown, thickened, not more than 0.2 mm.

ascending, veinless, somewhat flattened, loosely enclosing achenes, ellipsoid or obovoid, dull, apex obtuse or acute, papillose.

Achenes

not constricted, glossy.

Pistillate

scales pale red-brown with narrow red-brown marginand broad pale midvein.

scales red-brown to purple-brown, apex obtuse or acute, awnless.

2n

= 76, 77.

= 72–80.

Carex aquatilis var. substricta

Carex aquatilis

Phenology Fruiting Jul–Aug.
Habitat Marshes, shallow water along shores, usually in neutral or calcareous substrates
Elevation 0–1000 m (0–3300 ft)
Distribution
from FNA
CT; IA; IL; IN; KS; MA; ME; MI; MN; MO; ND; NE; NJ; NY; OH; PA; SD; VA; VT; WI; MB; NB; NS; ON; QC
from FNA
AK; AZ; CA; CO; CT; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NJ; NM; NV; NY; OH; OR; PA; SD; UT; VA; VT; WA; WI; WY; AB; BC; MB; NB; NS; NT; NU; ON; PE; QC; SK; YT; Europe
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[BONAP county map]
Discussion

Populations of Carex aquatilis var. substricta from Minnesota to Massachusetts tend to have very pale scales with a very broad, pale brown midvein, distinctly obovoid perigynia, and broad pistillate spikes that are rarely staminate at the apex. These populations tend to occur on neutral or calcareous substrates (fens) rather than the more acidic marshes favored by var. aquatilis. The two varieties intergrade where they come into contact.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

Carex aquatilis is circumboreal and variable; four extensively intergrading varieties are recognized in North America. The species is distinguished by amphistomic (epistomic in var. dives) papillose leaves, glabrous sheaths with a concave apex, perigynia that lack veins and are usually brown-spotted on the proximal half, and have glossy achenes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikes pendent or the terminal erect; proximal peduncle to 11 cm; perigynia ellipsoid; beak purple-brown, 0.3–0.4 mm; culms scabrous on angles.
var. dives
1. Spikes erect; proximal peduncle not more than 4 cm; perigynia usually obovoid; beak pale or purple-brown, not more than 0.2 mm; culms glabrous or scabrous on angles.
→ 2
2. Pistillate scales pale brown with narrow red-brown margins and broad pale midvein.
var. substricta
2. Pistillate scales red- or purple-brown and narrow pale midvein.
→ 3
3. Perigynia pale brown; staminate spikes usually 2–4.
var. aquatilis
3. Perigynia purple-brown on apical 1/2; staminate spikes usually 1–2.
var. minor
Source FNA vol. 23, p. 398. FNA vol. 23, p. 397.
Parent taxa Cyperaceae > Carex > sect. Phacocystis > Carex aquatilis Cyperaceae > Carex > sect. Phacocystis
Sibling taxa
C. aquatilis var. aquatilis, C. aquatilis var. dives, C. aquatilis var. minor
C. aboriginum, C. abrupta, C. abscondita, C. acidicola, C. acutiformis, C. adelostoma, C. adusta, C. aestivalis, C. aggregata, C. alata, C. albicans, C. albida, C. albolutescens, C. albonigra, C. albursina, C. alma, C. alopecoidea, C. amphibola, C. amplectens, C. amplifolia, C. angustata, C. annectens, C. anthoxanthea, C. aperta, C. appalachica, C. arapahoensis, C. arcta, C. arctata, C. arctiformis, C. arenaria, C. argyrantha, C. arkansana, C. assiniboinensis, C. atherodes, C. athrostachya, C. atlantica, C. atrata, C. atratiformis, C. atrofusca, C. atrosquama, C. aurea, C. aureolensis, C. austrina, C. austrocaroliniana, C. backii, C. baileyi, C. baltzellii, C. barbarae, C. barrattii, C. basiantha, C. bebbii, C. bella, C. bicknellii, C. bicolor, C. bigelowii, C. biltmoreana, C. blanda, C. bolanderi, C. bonanzensis, C. brainerdii, C. brevicaulis, C. brevior, C. breweri, C. bromoides, C. brunnescens, C. brysonii, C. bulbostylis, C. bullata, C. bushii, C. buxbaumii, C. calcifugens, C. californica, C. canescens, C. capillaris, C. capitata, C. careyana, C. caroliniana, C. caryophyllea, C. castanea, C. cephaloidea, C. cephalophora, C. chalciolepis, C. chapmanii, C. cherokeensis, C. chihuahuensis, C. chordorrhiza, C. circinata, C. collinsii, C. communis, C. comosa, C. complanata, C. concinna, C. concinnoides, C. congdonii, C. conjuncta, C. conoidea, C. constanceana, C. cordillerana, C. corrugata, C. crawei, C. crawfordii, C. crebriflora, C. crinita, C. cristatella, C. crus-corvi, C. cryptolepis, C. cumberlandensis, C. cumulata, C. curatorum, C. cusickii, C. dasycarpa, C. davisii, C. davyi, C. debilis, C. decomposita, C. deflexa, C. densa, C. deweyana, C. diandra, C. digitalis, C. diluta, C. disperma, C. distans, C. disticha, C. divisa, C. divulsa, C. douglasii, C. duriuscula, C. ebenea, C. eburnea, C. echinata, C. edwardsiana, C. egglestonii, C. eleusinoides, C. elliottii, C. elynoides, C. emoryi, C. endlichii, C. engelmannii, C. epapillosa, C. exilis, C. exsiccata, C. extensa, C. festucacea, C. feta, C. filifolia, C. fissa, C. fissuricola, C. flacca, C. flaccosperma, C. flava, C. floridana, C. foenea, C. folliculata, C. formosa, C. fracta, C. frankii, C. fuliginosa, C. garberi, C. geophila, C. geyeri, C. gholsonii, C. gigantea, C. glacialis, C. glareosa, C. glaucescens, C. glaucodea, C. globosa, C. gmelinii, C. godfreyi, C. gracilescens, C. gracilior, C. gracillima, C. granularis, C. gravida, C. grayi, C. grisea, C. gynandra, C. gynocrates, C. gynodynama, C. halliana, C. hallii, C. harfordii, C. hassei, C. haydeniana, C. haydenii, C. heleonastes, C. helleri, C. hendersonii, C. heteroneura, C. heterostachya, C. hirsutella, C. hirta, C. hirtifolia, C. hirtissima, C. hitchcockiana, C. holostoma, C. hoodii, C. hookeriana, C. hormathodes, C. hostiana, C. houghtoniana, C. hyalina, C. hyalinolepis, C. hystericina, C. idahoa, C. illota, C. impressinervia, C. incurviformis, C. infirminervia, C. inops, C. integra, C. interior, C. interrupta, C. intumescens, C. jamesii, C. jonesii, C. joorii, C. juniperorum, C. kobomugi, C. kraliana, C. krausei, C. lachenalii, C. lacustris, C. laeviconica, C. laeviculmis, C. laevivaginata, C. lapponica, C. lasiocarpa, C. latebracteata, C. lativena, C. laxa, C. laxiculmis, C. laxiflora, C. leavenworthii, C. lemmonii, C. lenticularis, C. leporinella, C. leptalea, C. leptonervia, C. leptopoda, C. limosa, C. livida, C. loliacea, C. lonchocarpa, C. longii, C. louisianica, C. lucorum, C. lupuliformis, C. lupulina, C. lurida, C. lutea, C. luzulifolia, C. luzulina, C. lyngbyei, C. mackenziei, C. macloviana, C. macrocephala, C. macrochaeta, C. magellanica, C. manhartii, C. marina, C. mariposana, C. maritima, C. mckittrickensis, C. meadii, C. media, C. melanostachya, C. membranacea, C. mendocinensis, C. merritt-fernaldii, C. mertensii, C. mesochorea, C. michauxiana, C. microchaeta, C. microdonta, C. microglochin, C. micropoda, C. microptera, C. misera, C. missouriensis, C. mitchelliana, C. molesta, C. molestiformis, C. muehlenbergii, C. multicaulis, C. multicostata, C. muricata, C. muriculata, C. muskingumensis, C. nardina, C. nebrascensis, C. nelsonii, C. nervina, C. neurophora, C. nigra, C. nigricans, C. nigromarginata, C. normalis, C. norvegica, C. nova, C. novae-angliae, C. nudata, C. obispoensis, C. obnupta, C. obtusata, C. occidentalis, C. oklahomensis, C. oligocarpa, C. oligosperma, C. opaca, C. oreocharis, C. ormostachya, C. oronensis, C. ouachitana, C. ovalis, C. oxylepis, C. ozarkana, C. pachystachya, C. paeninsulae, C. paleacea, C. pallescens, C. panicea, C. pansa, C. parallela, C. parryana, C. pauciflora, C. paysonis, C. peckii, C. pedunculata, C. pellita, C. pelocarpa, C. pendula, C. pensylvanica, C. perdentata, C. perglobosa, C. petasata, C. petricosa, C. phaeocephala, C. picta, C. pigra, C. pityophila, C. planispicata, C. planostachys, C. plantaginea, C. platyphylla, C. pluriflora, C. podocarpa, C. polymorpha, C. praeceptorum, C. praegracilis, C. prairea, C. prasina, C. praticola, C. preslii, C. projecta, C. proposita, C. pseudocyperus, C. pumila, C. purpurifera, C. radfordii, C. radiata, C. ramenskii, C. rariflora, C. raynoldsii, C. recta, C. reniformis, C. retroflexa, C. retrorsa, C. richardsonii, C. roanensis, C. rosea, C. rossii, C. rostrata, C. rotundata, C. rufina, C. rupestris, C. ruthii, C. sabulosa, C. salina, C. sartwelliana, C. sartwellii, C. saxatilis, C. saximontana, C. scabrata, C. scabriuscula, C. schottii, C. schweinitzii, C. scirpoidea, C. scoparia, C. scopulorum, C. senta, C. seorsa, C. serpenticola, C. serratodens, C. sheldonii, C. shinnersii, C. shortiana, C. siccata, C. silicea, C. simulata, C. socialis, C. sparganioides, C. specifica, C. spectabilis, C. specuicola, C. spicata, C. spissa, C. sprengelii, C. squarrosa, C. stenoptila, C. sterilis, C. stevenii, C. stipata, C. straminea, C. straminiformis, C. striata, C. striatula, C. stricta, C. styloflexa, C. stylosa, C. subbracteata, C. suberecta, C. subfusca, C. subnigricans, C. subspathacea, C. superata, C. supina, C. swanii, C. sychnocephala, C. sylvatica, C. tahoensis, C. tenax, C. tenera, C. tenuiflora, C. tetanica, C. tetrastachya, C. texensis, C. thornei, C. thurberi, C. timida, C. tincta, C. tompkinsii, C. tonsa, C. torreyi, C. torta, C. triangularis, C. tribuloides, C. trichocarpa, C. triquetra, C. trisperma, C. tuckermanii, C. tumulicola, C. turbinata, C. turgescens, C. typhina, C. umbellata, C. unilateralis, C. ursina, C. utriculata, C. vacillans, C. vaginata, C. vallicola, C. venusta, C. vernacula, C. verrucosa, C. vesicaria, C. vestita, C. vexans, C. virescens, C. viridula, C. vulpinoidea, C. whitneyi, C. wiegandii, C. willdenowii, C. williamsii, C. woodii, C. wootonii, C. xerantica
Subordinate taxa
C. aquatilis var. aquatilis, C. aquatilis var. dives, C. aquatilis var. minor, C. aquatilis var. substricta
Synonyms C. substricta
Name authority Kükenthal: in H. G. A. Engler, Pflanzenr. 38[IV,20]: 309. (1909) Wahlenberg: Kongl. Vetensk. Acad. Nya Handl. 24: 165. (1803)
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