Carex aquatilis var. aquatilis |
Carex aquatilis var. substricta |
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carex aquatique, water sedge |
calcareous water sedge, carex subétroit |
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Culms | 20–120 cm, angles glabrous. |
35–100 cm, angles glabrous or scabrous. |
Leaves | basal sheaths leafless, usually red-brown; blades amphistomic, 2.5–8 mm wide, papillose on both surfaces. |
basal sheaths leafless, usually red-brown; blades amphistomic, 5–8 mm wide, papillose on both surfaces. |
Inflorescences | peduncle of proximal spike 1 cm or longer; proximal bract slightly longer than inflorescence, 2–8 mm wide. |
peduncle of proximal spike not more than 4 cm; proximal bract longer than inflorescence, 4–8 mm wide. |
Spikes | erect; proximal 2–7 spikes pistillate, 1–10 cm × 3–7 mm, base cuneate; terminal 2–4 spikes usually staminate. |
erect; proximal 2–6 spikes pistillate, 4–6 cm × 4–7 mm, base cuneate or attenuate; terminal 1–3 spikes staminate. |
Perigynia | pale brown, ellipsoid or obovoid, 2–3.6 × 1.3–2.3 mm, apex rounded or obtuse; beak pale brown, thickened, not more than 0.2 mm. |
pale brown, veinless, obovoid, 2.6–3.2 × 1.5–2.2 mm, apex rounded; beak pale brown, thickened, not more than 0.2 mm. |
Pistillate | scales red-brown to purple-brown with narrow pale midvein. |
scales pale red-brown with narrow red-brown marginand broad pale midvein. |
2n | = 72, 74–77, 79–80. |
= 76, 77. |
Carex aquatilis var. aquatilis |
Carex aquatilis var. substricta |
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Phenology | Fruiting Jul–Aug. | Fruiting Jul–Aug. |
Habitat | Marshes, bogs, wet meadows, shallow water along shores, usually in acidic substrates | Marshes, shallow water along shores, usually in neutral or calcareous substrates |
Elevation | 0–3000 m (0–9800 ft) | 0–1000 m (0–3300 ft) |
Distribution |
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NM; NV; OR; UT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; SPM; Eurasia |
CT; IA; IL; IN; KS; MA; ME; MI; MN; MO; ND; NE; NJ; NY; OH; PA; SD; VA; VT; WI; MB; NB; NS; ON; QC |
Discussion | The circumboreal Carex aquatilis var. aquatilis is very common in wetlands of the northern and montane portions of North America. The plants are usually rhizomatous; in very wet or flooded sites they may form cespitose clumps. Carex aquatilis var. aquatilis is distinguished from the sympatric Carex stricta by the sheaths not ladder-fibrillose, obovoid and veinless perigynia that are rounded at apex, glossy achenes, and amphistomic leaves. It is often confused with C. emoryi, and distinguished by perigynia that lack veins, concave sheath apex, glossy achenes, and amphistomic leaves. In western North America, it is often sympatric with C. scopulorum and may be distinguished by the pale brown perigynia, glossy achenes, and amphistomic leaves. Carex aquatilis and C. scopulorum occasionally hybridize; hybrids have been named as C. ×sphacelata T. Holm and C. ×chionophila T. Holm. Carex aquatilis var. aquatilis overlaps and may intergrade with C. aquatilis var. dives along the Pacific Coast, with var. minor along the Arctic coast, and with var. substricta in the Great Lakes-New England region. Much of var. aquatilis in the southern part of its range has been called var. altior (L. H. Bailey) Rydberg. The type for var. altior is a specimen of C. emoryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Populations of Carex aquatilis var. substricta from Minnesota to Massachusetts tend to have very pale scales with a very broad, pale brown midvein, distinctly obovoid perigynia, and broad pistillate spikes that are rarely staminate at the apex. These populations tend to occur on neutral or calcareous substrates (fens) rather than the more acidic marshes favored by var. aquatilis. The two varieties intergrade where they come into contact. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 397. | FNA vol. 23, p. 398. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. acutinella, C. interimus, C. pachystoma, C. suksdorfii, C. variabilis | C. substricta |
Name authority | unknown | Kükenthal: in H. G. A. Engler, Pflanzenr. 38[IV,20]: 309. (1909) |
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