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compact-head thistle, Italian plumeless thistle, Italian thistle

chardon, plumeless thistle

Habit Annuals or biennials [perennials], 30–200(–400) cm, spiny, ± tomentose, sometimes glabrate.
Stems

erect, simple to much branched, (spiny-winged).

Leaves

basal and cauline; petiolate or sessile;

blade margins spiny dentate, often 1–2-pinnately lobed, faces glabrous or hairy, eglandular.

Receptacles

flat, epaleate, bearing setiform scales (“flattened bristles”).

Florets

several–many;

corollas white to pink or purple, ± bilateral, tubes long, slender, throats short, campanulate, abruptly expanded from tubes, lobes linear;

anther bases sharply short-tailed, apical appendages oblong;

style branches: fused portions with slightly, minutely puberulent, swollen basal nodes, distally papillate or glabrous, distinct portions very short.

Phyllaries

many in 7–10+ series, linear to broadly ovate, bases appressed, margins entire, apices ascending to spreading or reflexed, acute, spine-tipped.

Heads

discoid, borne singly or 2–20 in dense clusters or corymbiform arrays. (Peduncles naked or leafy-bracteate, spiny-winged or not winged.) Involucres cylindric to spheric.

Cypselae

ovoid, slightly compressed, faces smooth, glabrous, attachment scars slightly lateral;

pappi persistent or falling in rings, of many minutely barbed, basally connate bristles or setiform, minutely barbed scales (“minutely flattened bristles”).

x

= 8, 9, 10, 11, 13.

Carduus pycnocephalus

Carduus

Distribution
from FNA
AL; AR; CA; ID; MS; NY; OR; PA; SC; TX; Eurasia (Mediterranean region)
[WildflowerSearch map]
[BONAP county map]
from USDA
Eurasia; Africa [Introduced in North America]
[BONAP county map]
Discussion

Subspecies 2 (1 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 90 (5 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Phyllary appendages 2–7 mm wide, usually wider than appressed bases; peduncles often elongate, distally wingless; heads often nodding, usually borne singly or in leafy corymbiform arrays; involucres 20–70 mm diam
C. nutans
1. Phyllary appendages 0.5–1.5 mm wide, usually narrower than appressed bases; peduncles short, if present, usually winged throughout or wingless only near tip; heads erect, 1–many, often clustered at branch tips; involucres 7–30 mm diam
→ 2
2. Involucres spheric or hemispheric
→ 3
2. Involucres cylindric or narrowly ellipsoid
→ 4
3. Corollas 13–20 mm; heads 18–25 mm; involucres 14–20 mm; abaxial leaf faces glabrate except for long, curled, septate hairs along veins
C. acanthoides
3. Corollas 11–16 mm; heads 15–18 mm; involucres 12–17 mm; abaxial leaf faces sparsely to densely tomentose with fine, nonseptate hairs and often with curled, septate hairs along veins as well
C. crispus
4. Heads 1–5 at ends of branches; phyllaries not scarious-margined, ± persistently tomentose, distally scabrous on margins and faces
C. pycnocephalus
4. Heads 5–20 at ends of branches; phyllaries scarious-margined, glabrous or spar- ingly tomentose, distally ciliolate or glabrous
C. tenuiflorus
Source FNA vol. 19, p. 93. FNA vol. 19, p. 91. Author: David J. Keil.
Parent taxa Asteraceae > tribe Cardueae > Carduus Asteraceae > tribe Cardueae
Sibling taxa
C. acanthoides, C. crispus, C. nutans, C. tenuiflorus
Subordinate taxa
C. pycnocephalus subsp. pycnocephalus
C. acanthoides, C. crispus, C. nutans, C. pycnocephalus, C. tenuiflorus
Name authority Linnaeus: Sp. Pl. ed. 2, 2: 1151. (1763) Linnaeus: Sp. Pl. 2: 820. (1753): Gen. Pl. ed. 5, 358. (1754)
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