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sand-carpet, sand-mat

cardionema, sand-carpet, sandmat

Habit Herbs, perennial.
Taproots

often stout.

Stems

often forming dense mats, 5–30+ cm, often obscured by stipules and dense pubescence.

prostrate, much-branched proximally, terete to angular.

Leaves

stipules 1–8 mm, often nearly as long as leaves, lobes acuminate;

blade 2-grooved, 5–13 mm, apex finely spinose, glabrous.

opposite, not connate, connected by thickened margins from which stipules arise, sessile;

stipules 2 per node, silvery, lanceolate to ovate, margins entire to irregularly fimbriate, apex 2-lobed;

blade prominently 1-veined, needlelike, not succulent, apex spinose.

Inflorescences

axillary, 1–3-flowered clusters;

bracts paired, resembling stipules, smaller.

Flowers

densely woolly, hairs 0.5–1.5 mm;

sepals 5, margins fimbriate, awn 1.5–4 mm;

staminodes white, 0.3–0.5 mm.

sessile;

hypanthium cup-shaped, not abruptly expanded distally;

sepals 5, distinct, olive-green, oblong to obovate, 1.2–2.8 mm (excluding awns), indurate, herbaceous, margins whitish, scarious, apex spinose, prominently hooded, awned, hoods projecting inward and enclosing developing fruit, awns arising from near apex, widely divergent, very stout, spinose, 3 awns and associated sepals well developed, alternating with 2 reduced sepals;

nectaries not apparent;

stamens 3–5;

filaments distinct to base;

staminodes 5, adnate basally with alternating filaments, ovate-triangular, apex acuminate;

styles 2, distinct, subcapitate, 0.2 mm, glabrous proximally;

stigmas 2, subterminal, obscurely papillate (50x).

Seeds

1.4–1.6 mm.

tan with red spot at hilum, narrowly ovate, not compressed, smooth, marginal wing absent, appendage absent;

embryo peripheral, straight to slightly curved.

Utricles

apex apiculate.

ellipsoid to cylindric, teeth absent, indehiscent.

Cardionema ramosissimum

Cardionema

Phenology Flowering spring–early summer.
Habitat Sandy beaches, grassy bluffs, sand dunes
Elevation 0-200 m (0-700 ft)
Distribution
from FNA
CA; OR; WA; Mexico; South America (Chile, Ecuador, Peru)
[WildflowerSearch map]
[BONAP county map]
from USDA
Coastal w North America; South America (Chile, Ecuador, Peru)
[BONAP county map]
Discussion

The flowers are burlike in fruit with the utricle enclosed in the rigid, persistent calyx, the presumed unit of seed dispersal.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 6 (1 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 46. FNA vol. 5, p. 45. Author: Ronald L. Hartman.
Parent taxa Caryophyllaceae > subfam. Paronychioideae > Cardionema Caryophyllaceae > subfam. Paronychioideae
Subordinate taxa
C. ramosissimum
Synonyms Loeflingia ramosissima
Name authority (Weinmann) A. Nelson & J. F. Macbride: Bot. Gaz. 56: 473. (1913) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 3: 372. (1828)
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