Cardionema ramosissimum |
Cardionema |
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sand-carpet, sand-mat |
cardionema, sand-carpet, sandmat |
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Habit | Herbs, perennial. | |
Taproots | often stout. |
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Stems | often forming dense mats, 5–30+ cm, often obscured by stipules and dense pubescence. |
prostrate, much-branched proximally, terete to angular. |
Leaves | stipules 1–8 mm, often nearly as long as leaves, lobes acuminate; blade 2-grooved, 5–13 mm, apex finely spinose, glabrous. |
opposite, not connate, connected by thickened margins from which stipules arise, sessile; stipules 2 per node, silvery, lanceolate to ovate, margins entire to irregularly fimbriate, apex 2-lobed; blade prominently 1-veined, needlelike, not succulent, apex spinose. |
Inflorescences | axillary, 1–3-flowered clusters; bracts paired, resembling stipules, smaller. |
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Flowers | densely woolly, hairs 0.5–1.5 mm; sepals 5, margins fimbriate, awn 1.5–4 mm; staminodes white, 0.3–0.5 mm. |
sessile; hypanthium cup-shaped, not abruptly expanded distally; sepals 5, distinct, olive-green, oblong to obovate, 1.2–2.8 mm (excluding awns), indurate, herbaceous, margins whitish, scarious, apex spinose, prominently hooded, awned, hoods projecting inward and enclosing developing fruit, awns arising from near apex, widely divergent, very stout, spinose, 3 awns and associated sepals well developed, alternating with 2 reduced sepals; nectaries not apparent; stamens 3–5; filaments distinct to base; staminodes 5, adnate basally with alternating filaments, ovate-triangular, apex acuminate; styles 2, distinct, subcapitate, 0.2 mm, glabrous proximally; stigmas 2, subterminal, obscurely papillate (50x). |
Seeds | 1.4–1.6 mm. |
tan with red spot at hilum, narrowly ovate, not compressed, smooth, marginal wing absent, appendage absent; embryo peripheral, straight to slightly curved. |
Utricles | apex apiculate. |
ellipsoid to cylindric, teeth absent, indehiscent. |
Cardionema ramosissimum |
Cardionema |
|
Phenology | Flowering spring–early summer. | |
Habitat | Sandy beaches, grassy bluffs, sand dunes | |
Elevation | 0-200 m (0-700 ft) | |
Distribution |
CA; OR; WA; Mexico; South America (Chile, Ecuador, Peru)
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Coastal w North America; South America (Chile, Ecuador, Peru) |
Discussion | The flowers are burlike in fruit with the utricle enclosed in the rigid, persistent calyx, the presumed unit of seed dispersal. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 6 (1 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 46. | FNA vol. 5, p. 45. |
Parent taxa | Caryophyllaceae > subfam. Paronychioideae > Cardionema | Caryophyllaceae > subfam. Paronychioideae |
Subordinate taxa | ||
Synonyms | Loeflingia ramosissima | |
Name authority | (Weinmann) A. Nelson & J. F. Macbride: Bot. Gaz. 56: 473. (1913) | de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 3: 372. (1828) |
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