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sand mat, sand-carpet


often forming dense mats, 5–30+ cm, often obscured by stipules and dense pubescence.


stipules 1–8 mm, often nearly as long as leaves, lobes acuminate;

blade 2-grooved, 5–13 mm, apex finely spinose, glabrous.


densely woolly, hairs 0.5–1.5 mm;

sepals 5, margins fimbriate, awn 1.5–4 mm;

staminodes white, 0.3–0.5 mm.


1.4–1.6 mm.


apex apiculate.

Cardionema ramosissimum

Phenology Flowering spring–early summer.
Habitat Sandy beaches, grassy bluffs, sand dunes
Elevation 0-200 m [0-700 ft]

The flowers are burlike in fruit with the utricle enclosed in the rigid, persistent calyx, the presumed unit of seed dispersal.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

from FNA
CA; OR; WA; Mexico; South America (Chile, Ecuador, Peru)
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Parent taxa Caryophyllaceae > subfam. Paronychioideae > Cardionema
Synonyms Loeflingia ramosissima
Name authority (Weinmann) A. Nelson & J. F. Macbride: Bot. Gaz. 56: 473. (1913)
Source Flora of North America vol. 5, p. 46.
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