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purple bittercress

hairy bitter-cress, shotweed

Habit Perennials; (often cespitose); hirsute. Annuals; sparsely hirsute basally (at least on petiole of basal leaves), often glabrous distally.
Rhizomes

vertical, 1–3 mm diam., (not fleshy).

absent.

Stems

(simple or few to several from base), usually erect, unbranched, (0.3–)0.5–1.2(–1.5) dm.

erect, ascending, or decumbent, unbranched or branched basally and/or distally, (0.3–)1–3.5(–4.5) dm, (not flexuous).

Basal leaves

sometimes rosulate, usually 3 or 5-foliolate, rarely simple, (1.5–)2.5–7 cm, leaflets petiolulate or sessile;

petiole (1.2–)2–5 cm;

lateral lobes or leaflets sessile, blade similar to terminal, distinctly smaller;

terminal lobe or leaflet petiolulate (0.05–0.6 cm), blade reniform or suborbicular to broadly ovate, 0.3–1 cm × 4–15 mm, base subcordate to rounded, margins entire, repand, or obscurely 2-toothed.

(persistent to anthesis), rosulate, (5–)8–15(–22)-foliolate, (2–)3.5–15(–17) cm, leaflets petiolulate;

petiole 0.5–5 cm, (ciliate);

lateral leaflet blade oblong, ovate, obovate, or orbicular, smaller than terminal, margins entire, repand, crenate, or 3-lobed;

terminal leaflet (petiolule 0.2–1 cm), blade reniform or orbicular, 0.4–2 cm × 6–30 mm, margins entire, repand, dentate, or 3 or 5-lobed.

Cauline leaves

1–3, usually compound, rarely simple, petiolate, leaflets petiolulate or sessile;

petiole (0.2–)0.5–2 cm, base not auriculate;

similar to basal, smaller.

1–4(–6), compound as basal, petiolate, [(0.5–)1.2–5.5(–7) cm, including petiole], leaflets petiolulate;

blade base not auriculate;

leaflets similar to basal.

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, 2–3(–4) × 1.4–1.7 mm, lateral pair not saccate basally;

petals usually purple to pink, rarely white, obovate, 5–7(–9) × 3–4(–5) mm, (clawed, apex rounded);

filaments: median pairs 2.5–3.5 mm, lateral pair 1.5–3.5 mm;

anthers oblong, 0.6–0.9 mm.

sepals oblong, 1.5–2.5 × 0.3–0.7 mm, lateral pair not saccate;

petals (sometimes absent) white, spatulate, 2.5–4.5(–5) × 0.5–1.1 mm; (stamens usually 4, lateral pair often absent, rarely 5 or 6);

filaments 1.8–3 mm;

anthers ovate, 0.3–0.5 mm.

Fruiting pedicels

suberect, ascending, or divaricate, 5–12 mm, pubescent.

erect to ascending, (2–)3–10(–14) mm.

Fruits

linear, 1.5–2.5 cm × 1.5–1.8 mm;

ovules 10–14 per ovary;

style 1–2.5 mm.

linear, (torulose), (0.9–)1.5–2.5(–2.8) cm × (0.8–)1–1.4 mm, (often appressed to rachis);

ovules 14–40 per ovary;

style 0.1–0.6(–1) mm.

Seeds

brown, oblong to broadly ovoid, 1.7–2.1 × 1.4–1.7 mm.

light brown, oblong or subquadrate, 0.9–1.3(–1.5) × 0.6–0.9(–1.1) mm, (narrowly margined).

2n

= 96.

= 16.

Cardamine purpurea

Cardamine hirsuta

Phenology Flowering Jun–Aug. Flowering Feb–Jul.
Habitat Moist tundra, damp woods and ravines, alpine turf, river flats, peaty subarctic meadows, streamsides, moist slopes Roadsides, clearings, disturbed sites, slopes, cedar glades, mixed woods, meadows, fields, waste grounds, damp places, grassy areas
Elevation 600-1800 m (2000-5900 ft) 0-700 m (0-2300 ft)
Distribution
from FNA
AK; YT; e Asia (Russian Far East)
[BONAP county map]
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IL; KY; LA; MA; MD; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; ON; w Eurasia [Introduced in North America; introduced also in Central America, South America, e Asia (Japan), South Africa, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Herbarium specimens of Cardamine hirsuta have been misidentified as C. oligosperma.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 483. FNA vol. 7, p. 475.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. purpurea var. albiflora, C. purpurea var. albiflos, C. purpurea var. lactiflora
Name authority Chamisso & Schlechtendal: Linnaea 1: 20. (1826) Linnaeus: Sp. Pl. 2: 655. (1753)
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