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purple bittercress

angle bitter-cress, angle-leaf bitter-cress, seaside bitter-cress

Habit Perennials; (often cespitose); hirsute. Perennials; glabrous or sparsely pubescent.
Rhizomes

vertical, 1–3 mm diam., (not fleshy).

cylindrical, slender, to 2 mm diam.

Stems

(simple or few to several from base), usually erect, unbranched, (0.3–)0.5–1.2(–1.5) dm.

erect, unbranched, (1.5–)2.5–8.5(–10) dm, sparsely to densely hirsute at base.

Basal leaves

sometimes rosulate, usually 3 or 5-foliolate, rarely simple, (1.5–)2.5–7 cm, leaflets petiolulate or sessile;

petiole (1.2–)2–5 cm;

lateral lobes or leaflets sessile, blade similar to terminal, distinctly smaller;

terminal lobe or leaflet petiolulate (0.05–0.6 cm), blade reniform or suborbicular to broadly ovate, 0.3–1 cm × 4–15 mm, base subcordate to rounded, margins entire, repand, or obscurely 2-toothed.

Rhizomal leaves

3 (or 5)-foliolate, (4–)7–20(–22) cm, leaflets petiolulate or subsessile;

petiole (2–)4–12(–14) cm;

lateral leaflets subsessile, blade similar to terminal, larger or smaller in size;

terminal leaflet petiolulate [(0.3–)0.5–1.5 cm], blade ovate to broadly lanceolate, 1.5–7(–9) cm, base usually cuneate, rarely subreniform or obtuse, margins 3–5(–7)-lobed or -toothed, surfaces puberulent.

Cauline leaves

1–3, usually compound, rarely simple, petiolate, leaflets petiolulate or sessile;

petiole (0.2–)0.5–2 cm, base not auriculate;

similar to basal, smaller.

(3 or) 4–8, 3 (or 5)-foliolate, petiolate, leaflets petiolulate or sessile;

petiole 1–4 cm, base not auriculate;

lateral leaflets sessile, blade similar to terminal, smaller, margins usually dentate, rarely entire;

terminal leaflet sessile or petiolulate, blade broadly ovate to narrowly lanceolate, 2–7 cm × 6–40 mm, margins minutely pubescent.

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, 2–3(–4) × 1.4–1.7 mm, lateral pair not saccate basally;

petals usually purple to pink, rarely white, obovate, 5–7(–9) × 3–4(–5) mm, (clawed, apex rounded);

filaments: median pairs 2.5–3.5 mm, lateral pair 1.5–3.5 mm;

anthers oblong, 0.6–0.9 mm.

sepals oblong, 2.5–4 × 1.3–2 mm, lateral pair saccate basally;

petals usually white, rarely pinkish, obovate, 8–15 × 4–8 mm (clawed, apex rounded or emarginate);

filaments: median pairs 3.5–6 mm, lateral pair 2–3.5 mm;

anthers oblong, 0.8–1.2 mm.

Fruiting pedicels

suberect, ascending, or divaricate, 5–12 mm, pubescent.

ascending to divaricate, (9–)12–25 mm.

Fruits

linear, 1.5–2.5 cm × 1.5–1.8 mm;

ovules 10–14 per ovary;

style 1–2.5 mm.

linear, 1.5–3.2 cm × 1.4–2 mm;

ovules 10–16 per ovary;

style (0.5–)1–4 mm.

Seeds

brown, oblong to broadly ovoid, 1.7–2.1 × 1.4–1.7 mm.

dark brown, oblong, 1.8–2.3 × 1–1.2 mm.

2n

= 96.

= 40.

Cardamine purpurea

Cardamine angulata

Phenology Flowering Jun–Aug. Flowering Apr–Jun.
Habitat Moist tundra, damp woods and ravines, alpine turf, river flats, peaty subarctic meadows, streamsides, moist slopes Moist ground, stream banks, swampy or damp woods, thickets, wet meadows
Elevation 600-1800 m (2000-5900 ft) 0-900 m (0-3000 ft)
Distribution
from FNA
AK; YT; e Asia (Russian Far East)
[BONAP county map]
from FNA
AK; CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 7, p. 483. FNA vol. 7, p. 467.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. rotundifolia, C. rupicola, C. umbellata
C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. purpurea var. albiflora, C. purpurea var. albiflos, C. purpurea var. lactiflora C. angulata var. alba, C. angulata var. hirsuta, C. angulata var. pentaphylla, Dentaria grandiflora
Name authority Chamisso & Schlechtendal: Linnaea 1: 20. (1826) Hooker: Fl. Bor.-Amer. 1: 44. (1829)
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