Cardamine pachystigma |
Cardamine hirsuta |
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rock toothwort, serpentine bittercress |
hairy bitter-cress, shotweed |
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Habit | Perennials; glabrous. | Annuals; sparsely hirsute basally (at least on petiole of basal leaves), often glabrous distally. |
Rhizomes | ovoid to oblong, (3–)4–18 mm diam., (fleshy, deeply underground). |
absent. |
Stems | erect, unbranched, 1–3 dm. |
erect, ascending, or decumbent, unbranched or branched basally and/or distally, (0.3–)1–3.5(–4.5) dm, (not flexuous). |
Basal leaves | (persistent to anthesis), rosulate, (5–)8–15(–22)-foliolate, (2–)3.5–15(–17) cm, leaflets petiolulate; petiole 0.5–5 cm, (ciliate); lateral leaflet blade oblong, ovate, obovate, or orbicular, smaller than terminal, margins entire, repand, crenate, or 3-lobed; terminal leaflet (petiolule 0.2–1 cm), blade reniform or orbicular, 0.4–2 cm × 6–30 mm, margins entire, repand, dentate, or 3 or 5-lobed. |
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Rhizomal leaves | simple, 1.2–2.3 cm; petiole 6.7–19.5 cm; blade orbicular to reniform, or cordate to broadly ovate, 2.7–5.5 cm × 30–68 mm, base cordate to subtruncate, margins with obtuse teeth terminating in apiculae. |
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Cauline leaves | 2–5, (crowded distally), simple; petiole 0.3–2.6 cm, base not auriculate; blade similar to basal, 1.8–5.5 cm × 12–47mm, base cuneate or obtuse to subcordate, marginal teeth apiculate. |
1–4(–6), compound as basal, petiolate, [(0.5–)1.2–5.5(–7) cm, including petiole], leaflets petiolulate; blade base not auriculate; leaflets similar to basal. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 4–7 × 2–2.5 mm, lateral pair saccate basally; petals usually pink or purple, rarely white, obovate to oblanceolate, 14–18 × 5–7 mm, (claw to 9 mm, apex rounded); filaments: median pairs 7–8 mm, lateral pair 5.5–6.5 mm; anthers oblong, ca. 1.5 mm. |
sepals oblong, 1.5–2.5 × 0.3–0.7 mm, lateral pair not saccate; petals (sometimes absent) white, spatulate, 2.5–4.5(–5) × 0.5–1.1 mm; (stamens usually 4, lateral pair often absent, rarely 5 or 6); filaments 1.8–3 mm; anthers ovate, 0.3–0.5 mm. |
Fruiting pedicels | divaricate to ascending, 7–24 mm. |
erect to ascending, (2–)3–10(–14) mm. |
Fruits | linear, 3.2–5.4 cm × 2.2–4 mm; ovules 10–14 per ovary; style 4–7 mm. |
linear, (torulose), (0.9–)1.5–2.5(–2.8) cm × (0.8–)1–1.4 mm, (often appressed to rachis); ovules 14–40 per ovary; style 0.1–0.6(–1) mm. |
Seeds | brown, broadly ovoid, 2–2.8 × 1.8–2.5 mm. |
light brown, oblong or subquadrate, 0.9–1.3(–1.5) × 0.6–0.9(–1.1) mm, (narrowly margined). |
2n | = 16. |
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Cardamine pachystigma |
Cardamine hirsuta |
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Phenology | Flowering Mar–May. | Flowering Feb–Jul. |
Habitat | Forests, lava slides, talus, cliffs | Roadsides, clearings, disturbed sites, slopes, cedar glades, mixed woods, meadows, fields, waste grounds, damp places, grassy areas |
Elevation | 700-2900 m (2300-9500 ft) | 0-700 m (0-2300 ft) |
Distribution |
CA
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AL; AR; CA; CT; DC; DE; FL; GA; IL; KY; LA; MA; MD; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; ON; w Eurasia [Introduced in North America; introduced also in Central America, South America, e Asia (Japan), South Africa, Australia]
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Discussion | Cardamine pachystigma is known from the disjunct counties of Plumas, Tehama, and Tulare. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Herbarium specimens of Cardamine hirsuta have been misidentified as C. oligosperma. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 480. | FNA vol. 7, p. 475. |
Parent taxa | Brassicaceae > tribe Cardamineae > Cardamine | Brassicaceae > tribe Cardamineae > Cardamine |
Sibling taxa | ||
Synonyms | Dentaria californica var. pachystigma, C. californica var. pachystigma, Dentaria corymbosa var. grata, Dentaria pachystigma | |
Name authority | (S. Watson) Rollins: Harvard Pap. Bot. 1(4): 45. (1993) | Linnaeus: Sp. Pl. 2: 655. (1753) |
Web links |
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