The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

rock toothwort, serpentine bittercress

slender toothwort

Habit Perennials; glabrous. Perennials; usually glabrous, rarely sparsely pubescent.
Rhizomes

ovoid to oblong, (3–)4–18 mm diam., (fleshy, deeply underground).

(tuberiform, fragile) moniliform, segments fusiform, 3–6 mm diam. (fleshy).

Stems

erect, unbranched, 1–3 dm.

erect, unbranched, 1.2–3(–4) dm, glabrous or pubescent.

Rhizomal leaves

simple, 1.2–2.3 cm;

petiole 6.7–19.5 cm;

blade orbicular to reniform, or cordate to broadly ovate, 2.7–5.5 cm × 30–68 mm, base cordate to subtruncate, margins with obtuse teeth terminating in apiculae.

3-foliolate, to 24 cm, leaflets petiolulate or subsessile;

petiole (3–)5–12(–16) cm;

lateral leaflets subsessile or petiolulate (0.2–1 cm), blade similar to terminal leaflet or smaller;

terminal leaflet (petiolule) (0.2–)0.5–1.5(–2) cm, blade broadly ovate to rhombic-obovate, 1.5–6(–8) cm, base usually cuneate, rarely subtruncate, margins coarsely dentate to crenate or 3-lobed, surfaces puberulent or not.

Cauline leaves

2–5, (crowded distally), simple;

petiole 0.3–2.6 cm, base not auriculate;

blade similar to basal, 1.8–5.5 cm × 12–47mm, base cuneate or obtuse to subcordate, marginal teeth apiculate.

2 (or 3), 3-foliolate (usually alternate, rarely opposite, different in morphology from rhizomal), petiolate, leaflets petiolulate or sessile;

petiole 0.5–2 cm, base not auriculate;

lateral leaflets sessile, blade similar to terminal, smaller, margins usually dentate, rarely entire;

terminal leaflet sessile or petiolulate, blade narrowly lanceolate to narrowly oblong or oblong-lanceolate, 2–7 × 0.3–0.6 cm, margins minutely puberulent.

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, 4–7 × 2–2.5 mm, lateral pair saccate basally;

petals usually pink or purple, rarely white, obovate to oblanceolate, 14–18 × 5–7 mm, (claw to 9 mm, apex rounded);

filaments: median pairs 7–8 mm, lateral pair 5.5–6.5 mm;

anthers oblong, ca. 1.5 mm.

sepals oblong, 5–7.5 × 1–2 mm, lateral pair slightly saccate basally;

petals purple to pale pink, oblanceolate, 9–18 × 2–5 mm (clawed, apex rounded);

filaments: median pairs 5–10 mm, lateral pair 3.5–8 mm;

anthers linear, 1.5–3 mm.

Fruiting pedicels

divaricate to ascending, 7–24 mm.

ascending to divaricate, 15–40 mm.

Fruits

linear, 3.2–5.4 cm × 2.2–4 mm;

ovules 10–14 per ovary;

style 4–7 mm.

linear, 2.5–4 cm × 1.5–2.5 mm;

ovules 8–12 per ovary;

style (5–)7–11 mm.

Seeds

brown, broadly ovoid, 2–2.8 × 1.8–2.5 mm.

dark brown, oblong, 2–2.5 × 1–1.5 mm.

2n

= ca. 128.

Cardamine pachystigma

Cardamine angustata

Phenology Flowering Mar–May. Flowering Mar–May.
Habitat Forests, lava slides, talus, cliffs Moist woods, wooded ridges and bottomlands, floodplains, shady ravines, streambeds
Elevation 700-2900 m (2300-9500 ft) 300-1300 m (1000-4300 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; DC; DE; GA; IN; KY; MD; MS; NC; NJ; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Cardamine pachystigma is known from the disjunct counties of Plumas, Tehama, and Tulare.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cardamine heterophylla (Nuttall) Alph. Wood (1870), not Host (1797) is an illegitimate name, sometimes found in synonymy with C. angustata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 480. FNA vol. 7, p. 467.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms Dentaria californica var. pachystigma, C. californica var. pachystigma, Dentaria corymbosa var. grata, Dentaria pachystigma C. angustata var. ouachitana, Dentaria heterophylla
Name authority (S. Watson) Rollins: Harvard Pap. Bot. 1(4): 45. (1993) O. E. Schulz: Bot. Jahrb. Syst. 32: 349. (1903)
Web links