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cuckoo bitter-cress, lady smock, northern field bitter-cress

Brewer's bitter-cress, round bittercress

Habit Perennials; (cespitose); glabrous. Perennials; usually glabrous, rarely sparsely pubescent basally.
Rhizomes

absent.

cylindrical, slender (rarely slightly thickened at stem base), 1–3(–4) mm diam.

Stems

erect, unbranched or, rarely, branched, 0.5–1.6(–3.5) dm.

erect or decumbent basally, unbranched or branched, (0.6–)1.5–6(–7) dm.

Basal leaves

(7 or) 9–21-foliolate, (thick, veins impressed);

leaflets petiolulate or sessile;

lateral lobes or leaflets similar to terminal;

terminal lobe or leaflet blade orbicular, broadly ovate to lanceolate, base rounded to cuneate, margins usually entire.

Rhizomal leaves

absent.

Cauline leaves

2–4(–7), pinnatisect or pinnately compound, (7 or) 9–21-foliolate, (thick, veins impressed), petiolate, leaflets petiolulate or sessile;

petiole base not auriculate;

lobes or leaflets (of proximal leaves) (4–)7–10 each side of rachis, fewer distally, distal leaves with 4 or 5 lobes or leaflets each side of rachis;

terminal leaflet petiolulate or sessile, blade (or lobe) narrowly lanceolate to lanceolate, base cuneate, margins entire.

3–8(–11), 3 or 5-foliolate (rarely only terminal leaflet present), petiolate, leaflets petiolulate or subsessile;

petiole (0.7–)1–4(–6) cm, base not auriculate;

lateral leaflets (when present) subsessile or petiolulate (to 0.4 cm), blade similar to terminal, often much smaller and narrower than terminal;

terminal leaflet (petiolule 0.4–1.6 cm), blade usually ovate to orbicular, rarely subcordate, 1.5–4(–5) cm × 15–35(–50) mm, base truncate, rounded, or cordate, margins crenate, dentate, sinuate, or to 11-lobed.

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong or ovate, 3.6–4.4 mm, lateral pair saccate basally, (green with hyaline margins);

petals white-lilac, 9–12.3 × 4.8–6.8 mm, (clawed, apex rounded or emarginate);

filaments: median pairs 3.5–4.5 mm, lateral pair 2–3 mm;

anthers narrowly oblong, 0.9–1.4 mm.

sepals oblong, 2–3(–3.8) × 1–1.5 mm, lateral pair not saccate basally;

petals white, oblanceolate, 3.5–6(–7) × 1.5–2.5(–3) mm (not clawed, apex rounded or ± emarginate);

filaments: median pairs 2.5–3.5 mm, lateral pair 2–2.5 mm;

anthers oblong, 0.7–1 mm.

Fruiting pedicels

erect-ascending, 5–15 mm.

ascending to divaricate-ascending, (7–)10–20 mm.

Fruits

linear, 1–1.8 cm × ca. 1.5 mm;

ovules ca. 16 per ovary;

style ca. 1 mm, (stout).

linear, 1.5–3.5 cm × 1–1.5 mm;

ovules 14–28 per ovary;

style 0.2–1.5(–2.5) mm.

Seeds

brown, oblong, ca. 1.5 mm.

brown, oblong, 1–1.6 × 0.9–1.1 mm.

2n

= 56, 60, 64, 80–100.

Cardamine nymanii

Cardamine breweri

Phenology Flowering Jun–Aug. Flowering Jun–Jul.
Habitat Wet meadows, marshes, margins of ponds, along streams, seacoasts, swamps Stream banks, seepage, lakeshores, creeks, wet meadows, swamps, ponds
Elevation 1200-3000 m (3900-9800 ft)
Distribution
from FNA
AK; MB; NL; NT; NU; QC; YT; Greenland; n Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY; BC
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 7, p. 478. FNA vol. 7, p. 469.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. pratensis var. angustifolia C. breweri var. leibergii, C. breweri var. orbicularis, C. breweri var. oregana, C. callosicrenata, C. foliacea, C. hederifolia, C. leibergii, C. modocensis, C. orbicularis, C. oregana, C. vallicola, C. vallicola subsp. leibergii
Name authority Gandoger: Bull. Soc. Bot. France 72: 1043. (1925) S. Watson: Proc. Amer. Acad. Arts 10: 339. (1875)
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