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cuckoo bitter-cress, lady smock, northern field bitter-cress

angle bitter-cress, angle-leaf bitter-cress, seaside bitter-cress

Habit Perennials; (cespitose); glabrous. Perennials; glabrous or sparsely pubescent.
Rhizomes

absent.

cylindrical, slender, to 2 mm diam.

Stems

erect, unbranched or, rarely, branched, 0.5–1.6(–3.5) dm.

erect, unbranched, (1.5–)2.5–8.5(–10) dm, sparsely to densely hirsute at base.

Basal leaves

(7 or) 9–21-foliolate, (thick, veins impressed);

leaflets petiolulate or sessile;

lateral lobes or leaflets similar to terminal;

terminal lobe or leaflet blade orbicular, broadly ovate to lanceolate, base rounded to cuneate, margins usually entire.

Rhizomal leaves

3 (or 5)-foliolate, (4–)7–20(–22) cm, leaflets petiolulate or subsessile;

petiole (2–)4–12(–14) cm;

lateral leaflets subsessile, blade similar to terminal, larger or smaller in size;

terminal leaflet petiolulate [(0.3–)0.5–1.5 cm], blade ovate to broadly lanceolate, 1.5–7(–9) cm, base usually cuneate, rarely subreniform or obtuse, margins 3–5(–7)-lobed or -toothed, surfaces puberulent.

Cauline leaves

2–4(–7), pinnatisect or pinnately compound, (7 or) 9–21-foliolate, (thick, veins impressed), petiolate, leaflets petiolulate or sessile;

petiole base not auriculate;

lobes or leaflets (of proximal leaves) (4–)7–10 each side of rachis, fewer distally, distal leaves with 4 or 5 lobes or leaflets each side of rachis;

terminal leaflet petiolulate or sessile, blade (or lobe) narrowly lanceolate to lanceolate, base cuneate, margins entire.

(3 or) 4–8, 3 (or 5)-foliolate, petiolate, leaflets petiolulate or sessile;

petiole 1–4 cm, base not auriculate;

lateral leaflets sessile, blade similar to terminal, smaller, margins usually dentate, rarely entire;

terminal leaflet sessile or petiolulate, blade broadly ovate to narrowly lanceolate, 2–7 cm × 6–40 mm, margins minutely pubescent.

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong or ovate, 3.6–4.4 mm, lateral pair saccate basally, (green with hyaline margins);

petals white-lilac, 9–12.3 × 4.8–6.8 mm, (clawed, apex rounded or emarginate);

filaments: median pairs 3.5–4.5 mm, lateral pair 2–3 mm;

anthers narrowly oblong, 0.9–1.4 mm.

sepals oblong, 2.5–4 × 1.3–2 mm, lateral pair saccate basally;

petals usually white, rarely pinkish, obovate, 8–15 × 4–8 mm (clawed, apex rounded or emarginate);

filaments: median pairs 3.5–6 mm, lateral pair 2–3.5 mm;

anthers oblong, 0.8–1.2 mm.

Fruiting pedicels

erect-ascending, 5–15 mm.

ascending to divaricate, (9–)12–25 mm.

Fruits

linear, 1–1.8 cm × ca. 1.5 mm;

ovules ca. 16 per ovary;

style ca. 1 mm, (stout).

linear, 1.5–3.2 cm × 1.4–2 mm;

ovules 10–16 per ovary;

style (0.5–)1–4 mm.

Seeds

brown, oblong, ca. 1.5 mm.

dark brown, oblong, 1.8–2.3 × 1–1.2 mm.

2n

= 56, 60, 64, 80–100.

= 40.

Cardamine nymanii

Cardamine angulata

Phenology Flowering Jun–Aug. Flowering Apr–Jun.
Habitat Wet meadows, marshes, margins of ponds, along streams, seacoasts, swamps Moist ground, stream banks, swampy or damp woods, thickets, wet meadows
Elevation 0-900 m (0-3000 ft)
Distribution
from FNA
AK; MB; NL; NT; NU; QC; YT; Greenland; n Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 7, p. 478. FNA vol. 7, p. 467.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. pratensis var. angustifolia C. angulata var. alba, C. angulata var. hirsuta, C. angulata var. pentaphylla, Dentaria grandiflora
Name authority Gandoger: Bull. Soc. Bot. France 72: 1043. (1925) Hooker: Fl. Bor.-Amer. 1: 44. (1829)
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