Cardamine nymanii |
Cardamine angulata |
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cuckoo bitter-cress, lady smock, northern field bitter-cress |
angle bitter-cress, angle-leaf bitter-cress, seaside bitter-cress |
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Habit | Perennials; (cespitose); glabrous. | Perennials; glabrous or sparsely pubescent. |
Rhizomes | absent. |
cylindrical, slender, to 2 mm diam. |
Stems | erect, unbranched or, rarely, branched, 0.5–1.6(–3.5) dm. |
erect, unbranched, (1.5–)2.5–8.5(–10) dm, sparsely to densely hirsute at base. |
Basal leaves | (7 or) 9–21-foliolate, (thick, veins impressed); leaflets petiolulate or sessile; lateral lobes or leaflets similar to terminal; terminal lobe or leaflet blade orbicular, broadly ovate to lanceolate, base rounded to cuneate, margins usually entire. |
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Rhizomal leaves | 3 (or 5)-foliolate, (4–)7–20(–22) cm, leaflets petiolulate or subsessile; petiole (2–)4–12(–14) cm; lateral leaflets subsessile, blade similar to terminal, larger or smaller in size; terminal leaflet petiolulate [(0.3–)0.5–1.5 cm], blade ovate to broadly lanceolate, 1.5–7(–9) cm, base usually cuneate, rarely subreniform or obtuse, margins 3–5(–7)-lobed or -toothed, surfaces puberulent. |
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Cauline leaves | 2–4(–7), pinnatisect or pinnately compound, (7 or) 9–21-foliolate, (thick, veins impressed), petiolate, leaflets petiolulate or sessile; petiole base not auriculate; lobes or leaflets (of proximal leaves) (4–)7–10 each side of rachis, fewer distally, distal leaves with 4 or 5 lobes or leaflets each side of rachis; terminal leaflet petiolulate or sessile, blade (or lobe) narrowly lanceolate to lanceolate, base cuneate, margins entire. |
(3 or) 4–8, 3 (or 5)-foliolate, petiolate, leaflets petiolulate or sessile; petiole 1–4 cm, base not auriculate; lateral leaflets sessile, blade similar to terminal, smaller, margins usually dentate, rarely entire; terminal leaflet sessile or petiolulate, blade broadly ovate to narrowly lanceolate, 2–7 cm × 6–40 mm, margins minutely pubescent. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong or ovate, 3.6–4.4 mm, lateral pair saccate basally, (green with hyaline margins); petals white-lilac, 9–12.3 × 4.8–6.8 mm, (clawed, apex rounded or emarginate); filaments: median pairs 3.5–4.5 mm, lateral pair 2–3 mm; anthers narrowly oblong, 0.9–1.4 mm. |
sepals oblong, 2.5–4 × 1.3–2 mm, lateral pair saccate basally; petals usually white, rarely pinkish, obovate, 8–15 × 4–8 mm (clawed, apex rounded or emarginate); filaments: median pairs 3.5–6 mm, lateral pair 2–3.5 mm; anthers oblong, 0.8–1.2 mm. |
Fruiting pedicels | erect-ascending, 5–15 mm. |
ascending to divaricate, (9–)12–25 mm. |
Fruits | linear, 1–1.8 cm × ca. 1.5 mm; ovules ca. 16 per ovary; style ca. 1 mm, (stout). |
linear, 1.5–3.2 cm × 1.4–2 mm; ovules 10–16 per ovary; style (0.5–)1–4 mm. |
Seeds | brown, oblong, ca. 1.5 mm. |
dark brown, oblong, 1.8–2.3 × 1–1.2 mm. |
2n | = 56, 60, 64, 80–100. |
= 40. |
Cardamine nymanii |
Cardamine angulata |
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Phenology | Flowering Jun–Aug. | Flowering Apr–Jun. |
Habitat | Wet meadows, marshes, margins of ponds, along streams, seacoasts, swamps | Moist ground, stream banks, swampy or damp woods, thickets, wet meadows |
Elevation | 0-900 m (0-3000 ft) | |
Distribution |
AK; MB; NL; NT; NU; QC; YT; Greenland; n Eurasia
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AK; CA; OR; WA; BC
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Source | FNA vol. 7, p. 478. | FNA vol. 7, p. 467. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. pratensis var. angustifolia | C. angulata var. alba, C. angulata var. hirsuta, C. angulata var. pentaphylla, Dentaria grandiflora |
Name authority | Gandoger: Bull. Soc. Bot. France 72: 1043. (1925) | Hooker: Fl. Bor.-Amer. 1: 44. (1829) |
Web links |
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