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cuckoo bitter-cress, lady smock, northern field bitter-cress

Habit Perennials; (cespitose); glabrous. Annuals, biennials, or perennials; eglandular.
Rhizomes

absent.

Stems

erect, unbranched or, rarely, branched, 0.5–1.6(–3.5) dm.

Basal leaves

(7 or) 9–21-foliolate, (thick, veins impressed);

leaflets petiolulate or sessile;

lateral lobes or leaflets similar to terminal;

terminal lobe or leaflet blade orbicular, broadly ovate to lanceolate, base rounded to cuneate, margins usually entire.

Cauline leaves

2–4(–7), pinnatisect or pinnately compound, (7 or) 9–21-foliolate, (thick, veins impressed), petiolate, leaflets petiolulate or sessile;

petiole base not auriculate;

lobes or leaflets (of proximal leaves) (4–)7–10 each side of rachis, fewer distally, distal leaves with 4 or 5 lobes or leaflets each side of rachis;

terminal leaflet petiolulate or sessile, blade (or lobe) narrowly lanceolate to lanceolate, base cuneate, margins entire.

(rarely absent), usually petiolate, sometimes sessile;

blade (simple or compound), base auriculate or not, margins entire or dentate to pinnately lobed.

Trichomes

absent or simple.

Racemes

ebracteate.

usually ebracteate (Selenia bracteate throughout), often elongated in fruit.

Flowers

sepals oblong or ovate, 3.6–4.4 mm, lateral pair saccate basally, (green with hyaline margins);

petals white-lilac, 9–12.3 × 4.8–6.8 mm, (clawed, apex rounded or emarginate);

filaments: median pairs 3.5–4.5 mm, lateral pair 2–3 mm;

anthers narrowly oblong, 0.9–1.4 mm.

actinomorphic;

sepals erect to spreading or ascending, lateral pair seldom saccate basally;

petals white, yellow, pink, lilac, or purple, claw usually present, rarely absent, often distinct;

filaments unappendaged, not winged;

pollen 3-colpate.

Fruiting pedicels

erect-ascending, 5–15 mm.

Fruits

linear, 1–1.8 cm × ca. 1.5 mm;

ovules ca. 16 per ovary;

style ca. 1 mm, (stout).

silicles or siliques, dehiscent, unsegmented, terete, 4-angled, or latiseptate, rarely angustiseptate;

ovules 4–300[–numerous] per ovary;

style distinct or obsolete (absent in Subularia);

stigma usually entire.

Seeds

brown, oblong, ca. 1.5 mm.

biseriate or uniseriate;

cotyledons usually accumbent, sometimes incumbent.

2n

= 56, 60, 64, 80–100.

Cardamine nymanii

Brassicaceae tribe Cardamineae

Phenology Flowering Jun–Aug.
Habitat Wet meadows, marshes, margins of ponds, along streams, seacoasts, swamps
Distribution
from FNA
AK; MB; NL; NT; NU; QC; YT; Greenland; n Eurasia
[WildflowerSearch map]
[BONAP county map]
Nearly worldwide
Discussion

Genera 12, species ca. 335 (10 genera, 85 species in the flora).

The assignment of Subularia to Cardamineae is provisional and based solely on morphology.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 478. FNA vol. 7, p. 458.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Subordinate taxa
Synonyms C. pratensis var. angustifolia
Name authority Gandoger: Bull. Soc. Bot. France 72: 1043. (1925) Dumortier: Fl. Belg., 124. (1827)
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