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streambank bittercress

big western bitter-cress, western bitter-cress, western toothwort

Habit Perennials; glabrous throughout. Perennials; glabrous or hirsute.
Rhizomes

(relatively short), 2–3 mm diam., (covered with extensive root system).

(tuberiform, fragile), ovoid or globose at base of stem, 3–10 mm diam., (fleshy).

Stems

erect to ascending, unbranched or branched distally, 0.9–4 dm.

(simple from base), erect to ascending, (not flexuous), unbranched or branched distally, 1–5 dm, glabrous or pubescent proximally.

Basal leaves

not rosulate, pinnately compound, (3 or) 5 (or 7)-foliolate, 2–10 cm, leaflets petiolulate or subsessile;

petiole 0.5–6.5 cm;

lateral leaflets petiolulate or subsessile, blade similar to terminal, ovate, smaller, margins entire;

terminal leaflet (petiolule 0.03–0.18 cm), blade orbicular to broadly ovate or subcordate, 0.5–2 cm × 7–25 mm, base cordate to rounded, margins entire or repand, (surfaces glabrous).

Rhizomal leaves

usually 3-foliolate, rarely simple, 1–8 cm, leaflets petiolulate or subsessile;

petiole 0.5–5 cm;

lateral leaflets subsessile, often minute;

terminal leaflet (petiolule 0.4–2 cm), blade orbicular to broadly ovate, 0.5–3 cm × 5–25 mm, base rounded, margins entire, repand, or dentate.

Cauline leaves

5–10, petiolate;

middle leaves often simple, petiole 0.3–1.5 cm, blade rhombic to suborbicular or ovate, 1–3.5 cm × 6–22 mm, base obtuse to cuneate, margins entire, repand, or dentate;

distal ones with shorter petiole, blade smaller.

3–7, (3 or) 5 or 7-foliolate (middle ones 5 or 7-foliolate, smaller distally, becoming 3-foliolate), petiolate;

petiole 0.5–3 cm, base not auriculate;

lateral leaflets similar to terminal, smaller;

terminal leaflet blade obovate to oblanceolate, 0.5–2.6 cm × 3–13 mm, margins shallowly toothed, entire, or repand.

Racemes

ebracteate.

ebracteate.

Flowers

sepals (ascending), oblong, 1.5–2.2 × 0.7–1 mm, lateral pair not saccate basally;

petals (somewhat spreading), white, oblanceolate, 3.5–5 × 1.2–1.8 mm, (not clawed, apex rounded);

filaments: median pairs 2.5–3 mm, lateral pair 2–2.5 mm;

anthers ovate, ca. 0.2 mm.

sepals oblong, 1.7–2 × 1–1.2 mm, lateral pair not saccate basally;

petals white, oblanceolate, 4–6 × 1.5–2 mm, (not clawed);

filaments: median pairs 2–2.5 mm, lateral pair 1–1.5 mm;

anthers ovate, 0.3–0.5 mm.

Fruiting pedicels

divaricate, 9–17 mm.

divaricate-ascending, 7–18 mm.

Fruits

linear, 0.8–1.6 cm × 0.8–1 mm;

ovules 16–22 per ovary;

style 1.2–1.8 mm.

linear, (torulose), 1.5–3.3 cm × 1.7–2.2 mm; (valves glabrous or sparsely pubescent);

ovules 18–40 per ovary;

style 0.5–1.5 mm.

Seeds

brown, oblong to ovoid, 0.9–1.2 × 0.6–0.8 mm.

brown, ovoid, 1–1.6 × 1–1.2 mm.

2n

= 64.

Cardamine micranthera

Cardamine occidentalis

Phenology Flowering Apr–May. Flowering Apr–Jul.
Habitat Wet grounds along streams, seepage, gravelly sandbars, moist crevices Muddy grounds, lake margins, shallow streams, meadows
Elevation 150-1500 m (500-4900 ft)
Distribution
from FNA
NC
[BONAP county map]
from FNA
AK; CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Cardamine micranthera is known only from Stokes County. It is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 477. FNA vol. 7, p. 479.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. pratensis var. occidentalis, C. neglecta
Name authority Rollins: Castanea 5: 87. (1940) (S. Watson) Howell: Fl. N.W. Amer., 50. (1897)
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