The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

large toothwort

streambank bittercress

Habit Perennials; glabrous (except leaflet margins and, sometimes, rachis). Perennials; glabrous throughout.
Rhizomes

cylindrical, 3–6 mm diam., (distinctly constricted at intervals, non-uniform diam., fleshy, slightly fragile, with dentate leaf scars).

(relatively short), 2–3 mm diam., (covered with extensive root system).

Stems

erect, unbranched, 0.9–3(–4) dm.

erect to ascending, unbranched or branched distally, 0.9–4 dm.

Rhizomal leaves

3-foliolate, 7–20 cm, leaflets petiolulate or subsessile;

petiole 4–15 cm;

lateral leaflets subsessile or petiolulate, blade often similar to terminal, base often oblique;

terminal leaflet (petiolule 0.2–1(–1.7) cm), blade broadly ovate to oblong, 2–7.5 cm × 12–37 mm, base cuneate to obtuse, margins coarsely dentate to sharply incised, or deeply cleft into 2 or 3 lobes (lobes dentate or incised, margins puberulent).

usually 3-foliolate, rarely simple, 1–8 cm, leaflets petiolulate or subsessile;

petiole 0.5–5 cm;

lateral leaflets subsessile, often minute;

terminal leaflet (petiolule 0.4–2 cm), blade orbicular to broadly ovate, 0.5–3 cm × 5–25 mm, base rounded, margins entire, repand, or dentate.

Cauline leaves

2 or 3, 3-foliolate, (rarely subopposite), petiolate, leaflets petiolulate or subsessile;

petiole (0.5–)1–4(–6.5) cm, base not auriculate;

lateral and terminal leaflets similar to rhizomal, distalmost sometimes much smaller.

5–10, petiolate;

middle leaves often simple, petiole 0.3–1.5 cm, blade rhombic to suborbicular or ovate, 1–3.5 cm × 6–22 mm, base obtuse to cuneate, margins entire, repand, or dentate;

distal ones with shorter petiole, blade smaller.

Racemes

ebracteate.

ebracteate.

Flowers

sepals (erect to ascending), oblong, 5–7 × 2–3 mm, lateral pair slightly saccate basally;

petals white or pink, oblanceolate, 10–17 × 3–6 mm, (not clawed, apex rounded);

filaments: median pairs 4–8 mm, lateral pair 3–6.5 mm;

anthers linear, 1.7–2.7 mm.

sepals (ascending), oblong, 1.5–2.2 × 0.7–1 mm, lateral pair not saccate basally;

petals (somewhat spreading), white, oblanceolate, 3.5–5 × 1.2–1.8 mm, (not clawed, apex rounded);

filaments: median pairs 2.5–3 mm, lateral pair 2–2.5 mm;

anthers ovate, ca. 0.2 mm.

Fruiting pedicels

(flowering ones) horizontal to divaricate or deflexed, 7–20 mm.

divaricate, 9–17 mm.

Fruits

(undeveloped), linear-lanceolate, to 3 cm × 2 mm;

ovules 10–14 per ovary;

style 3.5–7 mm.

linear, 0.8–1.6 cm × 0.8–1 mm;

ovules 16–22 per ovary;

style 1.2–1.8 mm.

Seeds

not known.

brown, oblong to ovoid, 0.9–1.2 × 0.6–0.8 mm.

2n

= 120, 124, 132, 138, 156, 161, ca. 208.

Cardamine maxima

Cardamine micranthera

Phenology Flowering Apr–Jun. Flowering Apr–May.
Habitat Rich woods, shady ravines, ledges, moist alluvial bottoms, steep forested slopes, stream banks Wet grounds along streams, seepage, gravelly sandbars, moist crevices
Distribution
from FNA
CT; MA; ME; MI; NJ; NY; OH; PA; VT; NB; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
NC
[BONAP county map]
Discussion

Cardamine maxima has not been found with mature fruits and seeds and it has long been suspected to be a hybrid between C. concatenata and C. diphylla. Molecular studies (P. W. Sweeney and R. A. Price 2000) indicate that C. maxima is distinct from both those species. Although we hesitate to maintain it as a species, its wide distribution and morphological distinctness warrant its recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Cardamine micranthera is known only from Stokes County. It is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 477. FNA vol. 7, p. 477.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms Dentaria maxima, C. anomala, Dentaria anomala
Name authority (Nuttall) Alph. Wood: Amer. Bot. Fl., 38. (1870) Rollins: Castanea 5: 87. (1940)
Web links