Cardamine maxima |
Cardamine breweri |
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large toothwort |
Brewer's bitter-cress, round bittercress |
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Habit | Perennials; glabrous (except leaflet margins and, sometimes, rachis). | Perennials; usually glabrous, rarely sparsely pubescent basally. |
Rhizomes | cylindrical, 3–6 mm diam., (distinctly constricted at intervals, non-uniform diam., fleshy, slightly fragile, with dentate leaf scars). |
cylindrical, slender (rarely slightly thickened at stem base), 1–3(–4) mm diam. |
Stems | erect, unbranched, 0.9–3(–4) dm. |
erect or decumbent basally, unbranched or branched, (0.6–)1.5–6(–7) dm. |
Rhizomal leaves | 3-foliolate, 7–20 cm, leaflets petiolulate or subsessile; petiole 4–15 cm; lateral leaflets subsessile or petiolulate, blade often similar to terminal, base often oblique; terminal leaflet (petiolule 0.2–1(–1.7) cm), blade broadly ovate to oblong, 2–7.5 cm × 12–37 mm, base cuneate to obtuse, margins coarsely dentate to sharply incised, or deeply cleft into 2 or 3 lobes (lobes dentate or incised, margins puberulent). |
absent. |
Cauline leaves | 2 or 3, 3-foliolate, (rarely subopposite), petiolate, leaflets petiolulate or subsessile; petiole (0.5–)1–4(–6.5) cm, base not auriculate; lateral and terminal leaflets similar to rhizomal, distalmost sometimes much smaller. |
3–8(–11), 3 or 5-foliolate (rarely only terminal leaflet present), petiolate, leaflets petiolulate or subsessile; petiole (0.7–)1–4(–6) cm, base not auriculate; lateral leaflets (when present) subsessile or petiolulate (to 0.4 cm), blade similar to terminal, often much smaller and narrower than terminal; terminal leaflet (petiolule 0.4–1.6 cm), blade usually ovate to orbicular, rarely subcordate, 1.5–4(–5) cm × 15–35(–50) mm, base truncate, rounded, or cordate, margins crenate, dentate, sinuate, or to 11-lobed. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals (erect to ascending), oblong, 5–7 × 2–3 mm, lateral pair slightly saccate basally; petals white or pink, oblanceolate, 10–17 × 3–6 mm, (not clawed, apex rounded); filaments: median pairs 4–8 mm, lateral pair 3–6.5 mm; anthers linear, 1.7–2.7 mm. |
sepals oblong, 2–3(–3.8) × 1–1.5 mm, lateral pair not saccate basally; petals white, oblanceolate, 3.5–6(–7) × 1.5–2.5(–3) mm (not clawed, apex rounded or ± emarginate); filaments: median pairs 2.5–3.5 mm, lateral pair 2–2.5 mm; anthers oblong, 0.7–1 mm. |
Fruiting pedicels | (flowering ones) horizontal to divaricate or deflexed, 7–20 mm. |
ascending to divaricate-ascending, (7–)10–20 mm. |
Fruits | (undeveloped), linear-lanceolate, to 3 cm × 2 mm; ovules 10–14 per ovary; style 3.5–7 mm. |
linear, 1.5–3.5 cm × 1–1.5 mm; ovules 14–28 per ovary; style 0.2–1.5(–2.5) mm. |
Seeds | not known. |
brown, oblong, 1–1.6 × 0.9–1.1 mm. |
2n | = 120, 124, 132, 138, 156, 161, ca. 208. |
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Cardamine maxima |
Cardamine breweri |
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Phenology | Flowering Apr–Jun. | Flowering Jun–Jul. |
Habitat | Rich woods, shady ravines, ledges, moist alluvial bottoms, steep forested slopes, stream banks | Stream banks, seepage, lakeshores, creeks, wet meadows, swamps, ponds |
Elevation | 1200-3000 m (3900-9800 ft) | |
Distribution |
CT; MA; ME; MI; NJ; NY; OH; PA; VT; NB; ON; QC
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CA; CO; ID; MT; NV; OR; UT; WA; WY; BC
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Discussion | Cardamine maxima has not been found with mature fruits and seeds and it has long been suspected to be a hybrid between C. concatenata and C. diphylla. Molecular studies (P. W. Sweeney and R. A. Price 2000) indicate that C. maxima is distinct from both those species. Although we hesitate to maintain it as a species, its wide distribution and morphological distinctness warrant its recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 477. | FNA vol. 7, p. 469. |
Parent taxa | Brassicaceae > tribe Cardamineae > Cardamine | Brassicaceae > tribe Cardamineae > Cardamine |
Sibling taxa | ||
Synonyms | Dentaria maxima, C. anomala, Dentaria anomala | C. breweri var. leibergii, C. breweri var. orbicularis, C. breweri var. oregana, C. callosicrenata, C. foliacea, C. hederifolia, C. leibergii, C. modocensis, C. orbicularis, C. oregana, C. vallicola, C. vallicola subsp. leibergii |
Name authority | (Nuttall) Alph. Wood: Amer. Bot. Fl., 38. (1870) | S. Watson: Proc. Amer. Acad. Arts 10: 339. (1875) |
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