Cardamine longii
Cardamine flexuosa
long's bitter-cress
wavy bitter-cress, wood bitter-cress, woodland bitter-cress
cylindrical, slender, 0.8–1.5 mm diam.
absent.
prostrate to decumbent or erect, unbranched or branched, 0.5–4(–6) dm.
erect, ascending, or decumbent, branched or unbranched, (0.6–)1–5 dm, (slightly flexuous).
(often withered by anthesis), not rosulate, 5–15-foliolate, (2.7–)4–14(–19) cm, leaflets petiolulate;
petiole 0.7–5 cm, (ciliate or not);
lateral leaflet blade oblong, ovate, or elliptic, smaller than terminal, margins entire, repand, crenate, or 3 (or 5)-lobed;
terminal leaflet (petiolule 0.3–1.7 cm), blade reniform, broadly ovate, or suborbicular, 0.5–2.5 cm × 4–30 mm, margins repand, crenate, or 3 or 5-lobed.
absent.
3–10, usually simple, rarely 3 or 5-foliolate, petiolate;
petiolulate or sessile;
petiole 0.4–2.5 cm, base not auriculate;
lateral leaflets (when present) sessile or petiolulate (to 0.2 cm), blade similar to terminal, considerably smaller;
terminal leaflet blade orbicular to reniform or ovate to oblong, 0.4–3 cm × 3–20 mm, (somewhat fleshy), base subcordate to rounded or subtruncate, margins entire or repand, rarely undulate, (apex rounded).
3–15, 5–15-foliolate [leaves (2–)3.5–5.5(–7) cm, including petiole], petiolate, leaflets petiolulate;
petiole base not auriculate;
lateral leaflets similar to basal, (0.4–2.5 mm wide).
ebracteate.
ebracteate.
sepals ovate, 0.7–1.2 × 0.5–0.7 mm, lateral pair not saccate basally;
petals absent or rudimentary, to 0.7 mm;
filaments 0.5–0.8 mm;
anthers ovate, 0.1–0.2 mm.
sepals oblong, 1.5–2.5 × 0.7–1 mm, lateral pair not saccate basally;
petals white, spatulate, 2.5–4(–5) × 1–1.7 mm; (stamens rarely 4, lateral pair absent);
filaments 2–3 mm;
anthers ovate, 0.3–0.5 mm.
divaricate-ascending to spreading, 0.5–2(–4) mm.
divaricate or ascending, (5–)6–14(–17) mm.
narrowly oblong to linear, (3–)5–10(–15) × 0.8–1.2 mm;
ovules 6–22 per ovary;
style 0.2–0.5(–1) mm.
linear, (torulose), (0.8–)1.2–2.8 cm × 1–1.5 mm;
ovules 18–40 per ovary;
style 0.3–1(–1.5) mm.
light or yellowish brown, oblong or ovoid, 1–1.4 × 0.7–1 mm.
brown, oblong or subquadrate, 0.9–1.5 × 0.6–1 mm, (narrowly margined or not).
= 32.
Cardamine longii
Cardamine flexuosa
According to J. Lihová et al. (2006), the populations referred to Cardamine flexuosa in North America comprise two taxa of different polyploid origins and evolutionary histories: tetraploid C. flexuosa (2n = 32), native to Europe, and the octoploid taxon informally called “Asian C. flexuosa” (2n = 64), native to eastern Asia. For the latter, the name C. flexuosa subsp. debilis can be used. Nevertheless, these two taxa should be recognized at species level and the correct name for the Asian species should be sought. Based on available data, both taxa occupy the same habitats in North America, but the Asian taxon is much more widespread. The occurrence of European C. flexuosa was, until now, confirmed only for Washington, where both taxa have been recorded. More detailed studies of the North American distributions of both these weeds are needed.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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