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long's bitter-cress

Richardson's bittercress

Habit Perennials; glabrous throughout. Perennials; glabrous throughout.
Rhizomes

cylindrical, slender, 0.8–1.5 mm diam.

cylindrical, slender, 0.5–1.5 mm diam.

Stems

prostrate to decumbent or erect, unbranched or branched, 0.5–4(–6) dm.

erect, unbranched, (0.6–)1–2(–3) dm.

Rhizomal leaves

absent.

pinnately 5 or 7-foliolate, (2–)4–10 cm, (not fleshy), leaflets sessile;

petiole (1.5–)3–7(–9) cm;

lateral leaflets similar to terminal, blade sometimes smaller;

terminal leaflet blade linear to narrowly oblong or narrowly lanceolate, (0.5–)1–2.5(–3.3) cm × 0.5–4(–8) mm, base cuneate to attenuate, margins entire.

Cauline leaves

3–10, usually simple, rarely 3 or 5-foliolate, petiolate;

petiolulate or sessile;

petiole 0.4–2.5 cm, base not auriculate;

lateral leaflets (when present) sessile or petiolulate (to 0.2 cm), blade similar to terminal, considerably smaller;

terminal leaflet blade orbicular to reniform or ovate to oblong, 0.4–3 cm × 3–20 mm, (somewhat fleshy), base subcordate to rounded or subtruncate, margins entire or repand, rarely undulate, (apex rounded).

(1 or) 2 or 3, 5 or 7-foliolate, petiolate, leaflets sessile;

petiole (0.2–)0.5–1.2(–1.5) cm, base not auriculate;

lateral leaflets similar to terminal, blade smaller;

terminal leaflet blade narrowly lanceolate to linear, 1–3.2(–4) cm × 0.8–2.5(–3.5) mm, base cuneate to attenuate, margins entire (apex acute to acuminate).

Racemes

ebracteate.

ebracteate.

Flowers

sepals ovate, 0.7–1.2 × 0.5–0.7 mm, lateral pair not saccate basally;

petals absent or rudimentary, to 0.7 mm;

filaments 0.5–0.8 mm;

anthers ovate, 0.1–0.2 mm.

sepals ovate, 2.5–3.5 × 1.5–2.5 mm, lateral pair not saccate basally;

petals white, obovate, 5–9 × 2.5–5 mm (clawed, apex rounded);

filaments: median pairs 3–4.5 mm, lateral pair 1.5–2.5 mm;

anthers oblong, 1–1.5 mm.

Fruiting pedicels

divaricate-ascending to spreading, 0.5–2(–4) mm.

suberect to divaricate-ascending, (7–)10–25 mm.

Fruits

narrowly oblong to linear, (3–)5–10(–15) × 0.8–1.2 mm;

ovules 6–22 per ovary;

style 0.2–0.5(–1) mm.

linear, (1.5–)2–4 cm × 1.5–2 mm;

ovules 6–12 per ovary;

style (1–)1.5–2.5(–4) mm.

Seeds

light or yellowish brown, oblong or ovoid, 1–1.4 × 0.7–1 mm.

brown, oblong, 1.8–2.2 × 0.9–1.1 mm.

2n

= 28, 42.

Cardamine longii

Cardamine digitata

Phenology Flowering Jun–Sep. Flowering Jun–Aug.
Habitat Tidal marshes, mud flats, tidal shores of rivers, shallow water, swampy areas, shady rocky crevices covered at high tide Damp flats, stream banks, tundra, meadows, bluffs, hummocks, sandy beaches, slopes, mossy mats, sedge swales
Elevation 0-10 m (0-0 ft) 0-1400 m (0-4600 ft)
Distribution
from FNA
FL; MA; MD; ME; NC; NJ; NY; SC; VA
[BONAP county map]
from FNA
AK; NT; YT; ne Asia (Russian Far East, Siberia)
[BONAP county map]
Source FNA vol. 7, p. 476. FNA vol. 7, p. 472.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. digitata var. oxyphylla, C. hyperborea, C. hyperborea var. oxyphylla, C. richardsonii
Name authority Fernald: Rhodora 19: 91. (1917) Richardson: in J. Franklin, Narr. Journey Polar Sea, 743. (1823)
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