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long's bitter-cress

alpine bitter-cress

Habit Perennials; glabrous throughout. Perennials (somewhat cespitose, subscapose); glabrous throughout.
Rhizomes

cylindrical, slender, 0.8–1.5 mm diam.

absent (caudex usually with slender, rarely stout, rhizomelike branches).

Stems

prostrate to decumbent or erect, unbranched or branched, 0.5–4(–6) dm.

erect to ascending, branched (several), 0.1–0.8(–1.4) dm, glabrous.

Basal leaves

rosulate, simple, (0.6–)1.2–5(–7) cm;

petiole (3–)1–3.5(–5.5) cm;

blade usually ovate to oblong, rarely oblanceolate to obovate, (0.4–)0.8–1.7(–2.5) cm × (2–)5–10(–16) mm, base cuneate to obtuse, margins usually entire, rarely repand or obtusely small-toothed (apex obtuse).

Rhizomal leaves

absent.

Cauline leaves

3–10, usually simple, rarely 3 or 5-foliolate, petiolate;

petiolulate or sessile;

petiole 0.4–2.5 cm, base not auriculate;

lateral leaflets (when present) sessile or petiolulate (to 0.2 cm), blade similar to terminal, considerably smaller;

terminal leaflet blade orbicular to reniform or ovate to oblong, 0.4–3 cm × 3–20 mm, (somewhat fleshy), base subcordate to rounded or subtruncate, margins entire or repand, rarely undulate, (apex rounded).

0 or 1 (or 2), simple, shortly petiolate;

petiole base not auriculate;

blade similar to basal leaves, much smaller.

Racemes

ebracteate.

ebracteate.

Flowers

sepals ovate, 0.7–1.2 × 0.5–0.7 mm, lateral pair not saccate basally;

petals absent or rudimentary, to 0.7 mm;

filaments 0.5–0.8 mm;

anthers ovate, 0.1–0.2 mm.

sepals oblong, 2–3(–4) × 0.8–1.5(–2) mm, lateral pair not saccate basally;

petals white, oblanceolate, 4–5.5(–7) × 1.3–2(–2.5) mm (not clawed, apex rounded or emarginate);

filaments: median pairs 2.5–4 mm, lateral pair 2–3 mm;

anthers oblong, 0.5–0.9 mm.

Fruiting pedicels

divaricate-ascending to spreading, 0.5–2(–4) mm.

ascending to erect, 3–6(–8) mm.

Fruits

narrowly oblong to linear, (3–)5–10(–15) × 0.8–1.2 mm;

ovules 6–22 per ovary;

style 0.2–0.5(–1) mm.

linear, (0.8–)1.3–2.8(–3.7) cm × 1.3–2 mm;

ovules 8–18 per ovary;

style 0.5–3 mm.

Seeds

light or yellowish brown, oblong or ovoid, 1–1.4 × 0.7–1 mm.

brown, oblong, 1.5–2 × 0.9–1.2 mm.

2n

= 16.

Cardamine longii

Cardamine bellidifolia

Phenology Flowering Jun–Sep. Flowering Jun–Sep.
Habitat Tidal marshes, mud flats, tidal shores of rivers, shallow water, swampy areas, shady rocky crevices covered at high tide Mossy areas, tundra, marshes at stream headwaters, cliffs, talus slopes, barren chert slopes, moist rock crevices, rocky slopes, alpine streams, sandy beaches, moist rocky streambeds
Elevation 0-10 m (0-0 ft) 0-2300 m (0-7500 ft)
Distribution
from FNA
FL; MA; MD; ME; NC; NJ; NY; SC; VA
[BONAP county map]
from FNA
AK; CA; ME; NH; OR; WA; AB; BC; LB; NT; NU; QC; YT; Greenland; n Europe; Asia (Russian Far East, Siberia)
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 7, p. 476. FNA vol. 7, p. 468.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. bellidifolia var. beringensis, C. bellidifolia var. laxa, C. bellidifolia var. pachyphylla, C. bellidifolia var. pinnatifida
Name authority Fernald: Rhodora 19: 91. (1917) Linnaeus: Sp. Pl. 2: 654. (1753)
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