Cardamine impatiens |
Cardamine umbellata |
|
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narrow-leaf bitter-cress |
Siberian bittercress, umbel bitter-cress, umbellate bittercress |
|
Habit | Biennials or, rarely, annuals; usually glabrous, rarely sparsely pubescent basally. | Perennials; usually glabrous. |
Rhizomes | absent. |
often elongated, usually slender, rarely thickened, 1–2(–5) mm diam., (not fleshy). |
Stems | erect, (angled, sometimes flexuous), unbranched basally, usually branched distally, (1.2–)2–6.5(–9) dm. |
(simple or few to several from base), erect to ascending, (not flexuous), unbranched basally, sometimes branched distally, (0.3–)0.8–2.5(–3) dm. |
Basal leaves | (often withered by flowering), rosulate, similar to cauline, with fewer distal leaflets. |
(sometimes withered by anthesis), rosulate, pinnately compound, (3 or) 5 or 7 (or 9)-foliolate, 2–5(–9) cm, leaflets petiolulate or subsessile; lateral leaflets shortly petiolulate or subsessile, blade usually broadly ovate, rarely broadly obovate or orbicular, smaller than terminal, margins usually entire, rarely slightly 3 (or 5)-lobed or crenate; terminal leaflet subsessile, blade reniform or orbicular, 0.4–0.8(–1.2) cm × 5–9(–16) mm, margins entire or 3 (or 5)-lobed or crenate. |
Cauline leaves | 9–24, (9–)13–25-foliolate, petiolate, leaflets petiolulate; petiole 2–6 cm, base auriculate (auricles to 10 × 2.2 mm); lateral leaflets similar to terminal, blade often smaller; terminal leaflet (petiolule to 0.5 cm), blade orbicular, obovate, ovate, or lanceolate, 1–4(–5) cm × 5–17 mm, margins entire or 3–5(–9)-toothed or -lobed. |
3–5(–7), 3–7 (or 9)-foliolate, petiolate, leaflets subsessile or sessile; base not auriculate; lateral leaflets: blade narrowly obovate, oblanceolate to linear, margins similar to terminal; terminal leaflet blade narrowly obovate, ovate, oblanceolate, lanceolate, oblong, margins usually entire, sometimes 3-lobed or crenate. |
Racemes | ebracteate. |
ebracteate, (subumbellate, 2–8(–14)-flowered, rachis usually 3–20 mm). |
Flowers | sepals oblong, 1.2–2(–2.5) × 0.7–1(–1.2) mm, lateral pair not saccate basally; petals (rarely absent), white, oblanceolate, 1.5–4(–5) × 0.6–1.2 mm; filaments 2–3(–4) mm; anthers ovate, 0.3–0.5 mm. |
sepals (greenish or purplish), oblong, 1–2 × 0.5–1 mm, lateral pair not saccate basally; petals white, narrowly obovate, 2.5–5 × 1–3 mm. |
Fruiting pedicels | divaricate or ascending, 3.5–12(–15) mm. |
suberect to ascending, 3–8(–10) mm. |
Fruits | linear, (torulose), (1–)1.6–3(–3.5) cm × 0.9–1.5 mm; (valves glabrous or, rarely, pilose); ovules 10–30 per ovary; style 0.6–1.6(–2) mm. |
linear, (torulose), (1.3–)1.8–2.5(–3) cm × 0.8–1.5(–2) mm; (valves glabrous or sparsely pubescent); style 0.5–2 mm. |
Seeds | brown, oblong, 1.1–1.5 × 0.8–1 mm, (compressed, sometimes narrowly winged apically). |
brown, oblong, 1–1.5 × 0.8–1 mm. |
2n | = 16. |
= 32, 36, 48. |
Cardamine impatiens |
Cardamine umbellata |
|
Phenology | Flowering May–Jul. | Flowering Jun–Sep. |
Habitat | Streamsides, slopes, roadsides, fields, disturbed areas | Stream banks, tundra, alpine slopes, wetlands, damp, swampy and mossy areas, beach gravel and sand, alpine stream margins |
Elevation | 0-200 m (0-700 ft) | 0-1800 m (0-5900 ft) |
Distribution |
CT; KY; MI; MN; NH; OH; PA; VA; WV; ON; Eurasia [Introduced in North America; introduced also in South Africa]
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AK; WA; AB; BC; NT; YT; e Asia (Russian Far East)
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Discussion | Recent molecular data (J. Lihová et al. 2006) indicate that Cardamine umbellata, often treated as a variety of C. oligosperma, represents a distinct lineage more closely related to taxa from New Zealand; this does not exclude C. oligosperma as one of the possible parents of this polyploid. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 476. | FNA vol. 7, p. 484. |
Parent taxa | Brassicaceae > tribe Cardamineae > Cardamine | Brassicaceae > tribe Cardamineae > Cardamine |
Sibling taxa | ||
Synonyms | C. impatiens var. angustifolia | C. hirsuta subsp. kamtschatica, C. kamtschatica, C. oligosperma var. kamtschatica, C. sylvatica var. kamtschatica |
Name authority | Linnaeus: Sp. Pl. 2: 655. (1753) | Greene: Pittonia 3: 154. (1897) |
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