Cardamine impatiens |
Cardamine angustata |
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narrow-leaf bitter-cress |
slender toothwort |
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Habit | Biennials or, rarely, annuals; usually glabrous, rarely sparsely pubescent basally. | Perennials; usually glabrous, rarely sparsely pubescent. |
Rhizomes | absent. |
(tuberiform, fragile) moniliform, segments fusiform, 3–6 mm diam. (fleshy). |
Stems | erect, (angled, sometimes flexuous), unbranched basally, usually branched distally, (1.2–)2–6.5(–9) dm. |
erect, unbranched, 1.2–3(–4) dm, glabrous or pubescent. |
Basal leaves | (often withered by flowering), rosulate, similar to cauline, with fewer distal leaflets. |
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Rhizomal leaves | 3-foliolate, to 24 cm, leaflets petiolulate or subsessile; petiole (3–)5–12(–16) cm; lateral leaflets subsessile or petiolulate (0.2–1 cm), blade similar to terminal leaflet or smaller; terminal leaflet (petiolule) (0.2–)0.5–1.5(–2) cm, blade broadly ovate to rhombic-obovate, 1.5–6(–8) cm, base usually cuneate, rarely subtruncate, margins coarsely dentate to crenate or 3-lobed, surfaces puberulent or not. |
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Cauline leaves | 9–24, (9–)13–25-foliolate, petiolate, leaflets petiolulate; petiole 2–6 cm, base auriculate (auricles to 10 × 2.2 mm); lateral leaflets similar to terminal, blade often smaller; terminal leaflet (petiolule to 0.5 cm), blade orbicular, obovate, ovate, or lanceolate, 1–4(–5) cm × 5–17 mm, margins entire or 3–5(–9)-toothed or -lobed. |
2 (or 3), 3-foliolate (usually alternate, rarely opposite, different in morphology from rhizomal), petiolate, leaflets petiolulate or sessile; petiole 0.5–2 cm, base not auriculate; lateral leaflets sessile, blade similar to terminal, smaller, margins usually dentate, rarely entire; terminal leaflet sessile or petiolulate, blade narrowly lanceolate to narrowly oblong or oblong-lanceolate, 2–7 × 0.3–0.6 cm, margins minutely puberulent. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 1.2–2(–2.5) × 0.7–1(–1.2) mm, lateral pair not saccate basally; petals (rarely absent), white, oblanceolate, 1.5–4(–5) × 0.6–1.2 mm; filaments 2–3(–4) mm; anthers ovate, 0.3–0.5 mm. |
sepals oblong, 5–7.5 × 1–2 mm, lateral pair slightly saccate basally; petals purple to pale pink, oblanceolate, 9–18 × 2–5 mm (clawed, apex rounded); filaments: median pairs 5–10 mm, lateral pair 3.5–8 mm; anthers linear, 1.5–3 mm. |
Fruiting pedicels | divaricate or ascending, 3.5–12(–15) mm. |
ascending to divaricate, 15–40 mm. |
Fruits | linear, (torulose), (1–)1.6–3(–3.5) cm × 0.9–1.5 mm; (valves glabrous or, rarely, pilose); ovules 10–30 per ovary; style 0.6–1.6(–2) mm. |
linear, 2.5–4 cm × 1.5–2.5 mm; ovules 8–12 per ovary; style (5–)7–11 mm. |
Seeds | brown, oblong, 1.1–1.5 × 0.8–1 mm, (compressed, sometimes narrowly winged apically). |
dark brown, oblong, 2–2.5 × 1–1.5 mm. |
2n | = 16. |
= ca. 128. |
Cardamine impatiens |
Cardamine angustata |
|
Phenology | Flowering May–Jul. | Flowering Mar–May. |
Habitat | Streamsides, slopes, roadsides, fields, disturbed areas | Moist woods, wooded ridges and bottomlands, floodplains, shady ravines, streambeds |
Elevation | 0-200 m (0-700 ft) | 300-1300 m (1000-4300 ft) |
Distribution |
CT; KY; MI; MN; NH; OH; PA; VA; WV; ON; Eurasia [Introduced in North America; introduced also in South Africa]
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AL; AR; DC; DE; GA; IN; KY; MD; MS; NC; NJ; OH; PA; SC; TN; VA; WV
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Discussion | Cardamine heterophylla (Nuttall) Alph. Wood (1870), not Host (1797) is an illegitimate name, sometimes found in synonymy with C. angustata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 476. | FNA vol. 7, p. 467. |
Parent taxa | Brassicaceae > tribe Cardamineae > Cardamine | Brassicaceae > tribe Cardamineae > Cardamine |
Sibling taxa | ||
Synonyms | C. impatiens var. angustifolia | C. angustata var. ouachitana, Dentaria heterophylla |
Name authority | Linnaeus: Sp. Pl. 2: 655. (1753) | O. E. Schulz: Bot. Jahrb. Syst. 32: 349. (1903) |
Web links |