Cardamine holmgrenii |
Cardamine oligosperma |
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Holmgren's bitter-cress |
alpine bitter-cress, few-seed bitter-cress, Idaho bittercress, little western bitter-cress, Siberian bitter-cress |
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Habit | Perennials; glabrous throughout. | Annuals or biennials; usually sparsely hirsute (at least proximally), rarely glabrous. |
Rhizomes | slender, 0.5–1 mm diam. |
absent. |
Stems | erect to ascending, branched, 1.5–2 dm. |
(simple or few from base), erect to ascending, (not flexuous), unbranched or branched distally, (0.5–)0.8–3.2(–4.1) dm, usually pubescent throughout or proximally, rarely glabrous. |
Basal leaves | (persistent to anthesis), rosulate, pinnately compound, 5–9(–13)-foliolate, 2–8.5(–11) cm, leaflets petiolulate or subsessile; petiole 1–6 cm; lateral leaflets petiolulate or subsessile, blade obovate to oblanceolate, smaller than terminal, margins entire or crenate; terminal leaflet (petiolule 0.1–0.7 cm), blade usually orbicular to ovate, rarely oblong, 0.4–1.5(–2.3) cm × 3–10(–13) mm, base cordate to rounded, margins entire, or crenate-dentate to obscurely 3 or 5-lobed, (surfaces often hirsute, sometimes glabrous). |
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Cauline leaves | 3 or 4, 3- or 5-foliolate, petiolate, leaflets petiolulate; petiole 0.5–1.7 cm, base not auriculate; lateral leaflet blade linear to linear-lanceolate, 0.9–1.5 cm × 1–3 mm, margins entire; terminal leaflet (petiolule 0.3–0.5 cm), blade obovate to lanceolate (linear distally), 1.3–2.3 cm × 2–8 mm, base cuneate, margins entire or subapically 1- or 2-toothed. |
3–8, pinnately compound, similar to basal, smaller and fewer distally, petiolate, leaflets petiolulate or subsessile; petiole 0.5–2 cm, base not auriculate. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 1.2–1.5 × 0.6–0.8 mm, lateral pair not saccate basally; petals white, oblanceolate, 2–2.5 × 0.8–1 mm, (not clawed, apex rounded); filaments: median pairs 1.5–1.8 mm, lateral pair 1–1.2 mm; anthers ovate, 0.2–0.3 mm. |
sepals oblong, 1.3–1.8(–2) × 0.5–1 mm, lateral pair not saccate basally; petals white, narrowly spatulate to oblanceolate, 2.5–3.5 × 0.9–1.5 mm (not clawed); filaments: median pairs 1.7–2.5 mm, lateral pair 1.2–2 mm; anthers ovate, 0.3–0.5 mm. |
Fruiting pedicels | ascending to suberect, 5–14 mm. |
divaricate-ascending, (2–)3–9(–12) mm. |
Fruits | linear, 1–2 cm × ca. 1 mm; ovules 16–24 per ovary; style 0.5–0.7 mm. |
linear, (torulose), (1.3–)1.6–2.8 cm × 1–1.7 mm; (valves glabrous or sparsely pubescent); ovules 16–36(–42) per ovary; style 0.4–1(–1.5) mm. |
Seeds | brown, oblong, 1.2–1.5 × 0.8–1 mm. |
brown, oblong, 1–1.6 × 0.8–1.2 mm. |
Rhizomal | and basal leaves absent. |
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2n | = 16. |
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Cardamine holmgrenii |
Cardamine oligosperma |
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Phenology | Flowering Jul–Aug. | Flowering Mar–Jul. |
Habitat | Boggy slopes | Stream banks, shady banks, creek bottoms, lakeshores, meadows, moist areas, wooded slopes |
Elevation | 50-3300 m (200-10800 ft) | |
Distribution |
OR |
CA; CO; ID; MT; NV; OR; UT; WA; WY; BC; Mexico (Baja California)
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Discussion | Cardamine holmgrenii is known only from the type collection, from the Blue Mountains in Baker County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 475. | FNA vol. 7, p. 479. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. acuminata, C. hirsuta var. acuminata, C. hirsuta var. bracteata, C. hirsuta subsp. oligosperma, C. hirsuta var. parviflora, C. oligosperma var. bracteata, C. oligosperma var. lucens, C. oligosperma var. unijuga, C. unijuga | |
Name authority | Al-Shehbaz: Harvard Pap. Bot. 11: 275. (2007) | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 85. (1838) |
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