Cardamine hirsuta |
Cardamine angustata |
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hairy bitter-cress, shotweed |
slender toothwort |
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Habit | Annuals; sparsely hirsute basally (at least on petiole of basal leaves), often glabrous distally. | Perennials; usually glabrous, rarely sparsely pubescent. |
Rhizomes | absent. |
(tuberiform, fragile) moniliform, segments fusiform, 3–6 mm diam. (fleshy). |
Stems | erect, ascending, or decumbent, unbranched or branched basally and/or distally, (0.3–)1–3.5(–4.5) dm, (not flexuous). |
erect, unbranched, 1.2–3(–4) dm, glabrous or pubescent. |
Basal leaves | (persistent to anthesis), rosulate, (5–)8–15(–22)-foliolate, (2–)3.5–15(–17) cm, leaflets petiolulate; petiole 0.5–5 cm, (ciliate); lateral leaflet blade oblong, ovate, obovate, or orbicular, smaller than terminal, margins entire, repand, crenate, or 3-lobed; terminal leaflet (petiolule 0.2–1 cm), blade reniform or orbicular, 0.4–2 cm × 6–30 mm, margins entire, repand, dentate, or 3 or 5-lobed. |
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Rhizomal leaves | 3-foliolate, to 24 cm, leaflets petiolulate or subsessile; petiole (3–)5–12(–16) cm; lateral leaflets subsessile or petiolulate (0.2–1 cm), blade similar to terminal leaflet or smaller; terminal leaflet (petiolule) (0.2–)0.5–1.5(–2) cm, blade broadly ovate to rhombic-obovate, 1.5–6(–8) cm, base usually cuneate, rarely subtruncate, margins coarsely dentate to crenate or 3-lobed, surfaces puberulent or not. |
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Cauline leaves | 1–4(–6), compound as basal, petiolate, [(0.5–)1.2–5.5(–7) cm, including petiole], leaflets petiolulate; blade base not auriculate; leaflets similar to basal. |
2 (or 3), 3-foliolate (usually alternate, rarely opposite, different in morphology from rhizomal), petiolate, leaflets petiolulate or sessile; petiole 0.5–2 cm, base not auriculate; lateral leaflets sessile, blade similar to terminal, smaller, margins usually dentate, rarely entire; terminal leaflet sessile or petiolulate, blade narrowly lanceolate to narrowly oblong or oblong-lanceolate, 2–7 × 0.3–0.6 cm, margins minutely puberulent. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 1.5–2.5 × 0.3–0.7 mm, lateral pair not saccate; petals (sometimes absent) white, spatulate, 2.5–4.5(–5) × 0.5–1.1 mm; (stamens usually 4, lateral pair often absent, rarely 5 or 6); filaments 1.8–3 mm; anthers ovate, 0.3–0.5 mm. |
sepals oblong, 5–7.5 × 1–2 mm, lateral pair slightly saccate basally; petals purple to pale pink, oblanceolate, 9–18 × 2–5 mm (clawed, apex rounded); filaments: median pairs 5–10 mm, lateral pair 3.5–8 mm; anthers linear, 1.5–3 mm. |
Fruiting pedicels | erect to ascending, (2–)3–10(–14) mm. |
ascending to divaricate, 15–40 mm. |
Fruits | linear, (torulose), (0.9–)1.5–2.5(–2.8) cm × (0.8–)1–1.4 mm, (often appressed to rachis); ovules 14–40 per ovary; style 0.1–0.6(–1) mm. |
linear, 2.5–4 cm × 1.5–2.5 mm; ovules 8–12 per ovary; style (5–)7–11 mm. |
Seeds | light brown, oblong or subquadrate, 0.9–1.3(–1.5) × 0.6–0.9(–1.1) mm, (narrowly margined). |
dark brown, oblong, 2–2.5 × 1–1.5 mm. |
2n | = 16. |
= ca. 128. |
Cardamine hirsuta |
Cardamine angustata |
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Phenology | Flowering Feb–Jul. | Flowering Mar–May. |
Habitat | Roadsides, clearings, disturbed sites, slopes, cedar glades, mixed woods, meadows, fields, waste grounds, damp places, grassy areas | Moist woods, wooded ridges and bottomlands, floodplains, shady ravines, streambeds |
Elevation | 0-700 m (0-2300 ft) | 300-1300 m (1000-4300 ft) |
Distribution |
AL; AR; CA; CT; DC; DE; FL; GA; IL; KY; LA; MA; MD; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; ON; w Eurasia [Introduced in North America; introduced also in Central America, South America, e Asia (Japan), South Africa, Australia]
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AL; AR; DC; DE; GA; IN; KY; MD; MS; NC; NJ; OH; PA; SC; TN; VA; WV
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Discussion | Herbarium specimens of Cardamine hirsuta have been misidentified as C. oligosperma. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cardamine heterophylla (Nuttall) Alph. Wood (1870), not Host (1797) is an illegitimate name, sometimes found in synonymy with C. angustata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 475. | FNA vol. 7, p. 467. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. angustata var. ouachitana, Dentaria heterophylla | |
Name authority | Linnaeus: Sp. Pl. 2: 655. (1753) | O. E. Schulz: Bot. Jahrb. Syst. 32: 349. (1903) |
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