Cardamine flexuosa |
Cardamine rupicola |
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wavy bitter-cress, wood bitter-cress, woodland bitter-cress |
cliff bittercress |
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Habit | Annuals or biennials; sparsely to densely hirsute basally or throughout, or glabrous. | Perennials; glabrous throughout. |
Rhizomes | absent. |
cylindrical, slender, 1–2 mm diam. |
Stems | erect, ascending, or decumbent, branched or unbranched, (0.6–)1–5 dm, (slightly flexuous). |
erect or decumbent at base, unbranched, 0.6–2 dm. |
Basal leaves | (often withered by anthesis), not rosulate, 5–15-foliolate, (2.7–)4–14(–19) cm, leaflets petiolulate; petiole 0.7–5 cm, (ciliate or not); lateral leaflet blade oblong, ovate, or elliptic, smaller than terminal, margins entire, repand, crenate, or 3 (or 5)-lobed; terminal leaflet (petiolule 0.3–1.7 cm), blade reniform, broadly ovate, or suborbicular, 0.5–2.5 cm × 4–30 mm, margins repand, crenate, or 3 or 5-lobed. |
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Rhizomal leaves | palmately or subpalmately compound, 3 or 5 (or 7)-foliolate, 5.5–17(–22) cm, (fleshy), petiolate, leaflets petiolulate or subsessile; petiole 4–14(–17) cm; lateral leaflets subsessile, blade similar to terminal, sometimes smaller; terminal leaflet (petiolule 0.1–0.5 cm), blade ovate to lanceolate or elliptic-oblong, 1–3 cm × 6–17 mm, base cuneate or obtuse, margins entire, (apiculate). |
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Cauline leaves | 3–15, 5–15-foliolate [leaves (2–)3.5–5.5(–7) cm, including petiole], petiolate, leaflets petiolulate; petiole base not auriculate; lateral leaflets similar to basal, (0.4–2.5 mm wide). |
2 or 3, 3 or 5-foliolate, petiolate, leaflets petiolulate or sessile; petiole 0.7–4(–8) cm, base not auriculate; lateral leaflets sessile, blade similar to terminal, smaller; terminal leaflet petiolulate (0.1–0.5 cm), blade elliptic to oblong, or ovate, 1.2–3.5 cm × 4–25 mm, margins entire. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 1.5–2.5 × 0.7–1 mm, lateral pair not saccate basally; petals white, spatulate, 2.5–4(–5) × 1–1.7 mm; (stamens rarely 4, lateral pair absent); filaments 2–3 mm; anthers ovate, 0.3–0.5 mm. |
sepals oblong, 3–5 × 1.5–2 mm, lateral pair saccate basally; petals white, obovate, 8–14 × 4–7 mm, (short-clawed, apex rounded or subemarginate); filaments: median pairs 4–5 mm, lateral pair 2.5–3.5 mm; anthers oblong, 1–1.2 mm. |
Fruiting pedicels | divaricate or ascending, (5–)6–14(–17) mm. |
ascending to divaricate, 6–18 mm. |
Fruits | linear, (torulose), (0.8–)1.2–2.8 cm × 1–1.5 mm; ovules 18–40 per ovary; style 0.3–1(–1.5) mm. |
linear, 2–4 cm × 1.5–2.2 mm; ovules 10–14 per ovary; style 1–5 mm. |
Seeds | brown, oblong or subquadrate, 0.9–1.5 × 0.6–1 mm, (narrowly margined or not). |
brown, oblong, 1.8–2.2 × 1.2–1.5 mm. |
2n | = 32. |
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Cardamine flexuosa |
Cardamine rupicola |
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Phenology | Flowering Apr–Jul. | Flowering Jul–Aug. |
Habitat | Disturbed areas, fields, nurseries, plantations, gardens, flower beds, lawns, roadsides | Limestone talus slopes, loose limey shale, moist banks |
Elevation | 0-1100 m (0-3600 ft) | 2200-2700 m (7200-8900 ft) |
Distribution |
AL; CA; FL; GA; IL; IN; LA; MD; MI; NC; NY; OH; OR; RI; TX; VA; WA; BC; NF; ON; Europe; e Asia [Introduced in North America; introduced also in Mexico, Central America, South America, Australia]
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MT |
Discussion | According to J. Lihová et al. (2006), the populations referred to Cardamine flexuosa in North America comprise two taxa of different polyploid origins and evolutionary histories: tetraploid C. flexuosa (2n = 32), native to Europe, and the octoploid taxon informally called “Asian C. flexuosa” (2n = 64), native to eastern Asia. For the latter, the name C. flexuosa subsp. debilis can be used. Nevertheless, these two taxa should be recognized at species level and the correct name for the Asian species should be sought. Based on available data, both taxa occupy the same habitats in North America, but the Asian taxon is much more widespread. The occurrence of European C. flexuosa was, until now, confirmed only for Washington, where both taxa have been recorded. More detailed studies of the North American distributions of both these weeds are needed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cardamine rupicola is known from Flathead, Lewis and Clarke, and Missoula counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 474. | FNA vol. 7, p. 483. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. flexuosa subsp. debilis, C. flexuosa var. debilis, C. hirsuta subsp. flexuosa, C. scutata subsp. flexuosa | C. californica var. rupicola, Dentaria rupicola |
Name authority | Withering: Arr. Brit. Pl. ed. 3, 3: 578. (1796) | (O. E. Schulz) C. L. Hitchcock: in C. L. Hitchcock et al., Vasc. Pl. Pacif. N.W. 2: 474. (1964) |
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