Cardamine flexuosa |
Cardamine constancei |
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wavy bitter-cress, wood bitter-cress, woodland bitter-cress |
Constance's bittercress |
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Habit | Annuals or biennials; sparsely to densely hirsute basally or throughout, or glabrous. | Perennials; glabrous or sparsely hirsute. |
Rhizomes | absent. |
cylindrical, slender, to 2 mm diam. |
Stems | erect, ascending, or decumbent, branched or unbranched, (0.6–)1–5 dm, (slightly flexuous). |
erect, unbranched, 1.5–5 dm, glabrous or sparsely hirsute basally. |
Basal leaves | (often withered by anthesis), not rosulate, 5–15-foliolate, (2.7–)4–14(–19) cm, leaflets petiolulate; petiole 0.7–5 cm, (ciliate or not); lateral leaflet blade oblong, ovate, or elliptic, smaller than terminal, margins entire, repand, crenate, or 3 (or 5)-lobed; terminal leaflet (petiolule 0.3–1.7 cm), blade reniform, broadly ovate, or suborbicular, 0.5–2.5 cm × 4–30 mm, margins repand, crenate, or 3 or 5-lobed. |
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Rhizomal leaves | absent. |
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Cauline leaves | 3–15, 5–15-foliolate [leaves (2–)3.5–5.5(–7) cm, including petiole], petiolate, leaflets petiolulate; petiole base not auriculate; lateral leaflets similar to basal, (0.4–2.5 mm wide). |
4–7, simple, (crowded distally), petiolate; petiole (0.5–)1–4(–5) cm, base not auriculate; blade usually broadly ovate or ovate-lanceolate to ovate-elliptic, rarely obovate, 5–13 cm × 20–65 mm, base cuneate, margins coarsely serrate (with apiculae), or repand to undulate (margins minutely pubescent, apex acute). |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 1.5–2.5 × 0.7–1 mm, lateral pair not saccate basally; petals white, spatulate, 2.5–4(–5) × 1–1.7 mm; (stamens rarely 4, lateral pair absent); filaments 2–3 mm; anthers ovate, 0.3–0.5 mm. |
sepals oblong, 6–8 × 2–2.5 mm, lateral pair slightly saccate basally; petals pink, oblanceolate, 15–28 × 5–8 mm, (claw to 10 mm, apex rounded); filaments: median pairs 8–10 mm, lateral pair 4–6 mm; anthers linear, 2–3 mm. |
Fruiting pedicels | divaricate or ascending, (5–)6–14(–17) mm. |
divaricate-ascending to suberect, 10–22 mm. |
Fruits | linear, (torulose), (0.8–)1.2–2.8 cm × 1–1.5 mm; ovules 18–40 per ovary; style 0.3–1(–1.5) mm. |
linear, 2.5–3.5(–5) cm × 1.9–2.1 mm; ovules 12–16 per ovary; style 2–3.5 mm. |
Seeds | brown, oblong or subquadrate, 0.9–1.5 × 0.6–1 mm, (narrowly margined or not). |
brown, oblong, 2–2.5 × 1.2–1.5 mm. |
2n | = 32. |
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Cardamine flexuosa |
Cardamine constancei |
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Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Disturbed areas, fields, nurseries, plantations, gardens, flower beds, lawns, roadsides | Moist cliffs, wooded creek bottoms, shaded draws, hillsides, moist woods, mixed coniferous forests, granitic soils |
Elevation | 0-1100 m (0-3600 ft) | 400-600 m (1300-2000 ft) |
Distribution |
AL; CA; FL; GA; IL; IN; LA; MD; MI; NC; NY; OH; OR; RI; TX; VA; WA; BC; NF; ON; Europe; e Asia [Introduced in North America; introduced also in Mexico, Central America, South America, Australia]
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ID
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Discussion | According to J. Lihová et al. (2006), the populations referred to Cardamine flexuosa in North America comprise two taxa of different polyploid origins and evolutionary histories: tetraploid C. flexuosa (2n = 32), native to Europe, and the octoploid taxon informally called “Asian C. flexuosa” (2n = 64), native to eastern Asia. For the latter, the name C. flexuosa subsp. debilis can be used. Nevertheless, these two taxa should be recognized at species level and the correct name for the Asian species should be sought. Based on available data, both taxa occupy the same habitats in North America, but the Asian taxon is much more widespread. The occurrence of European C. flexuosa was, until now, confirmed only for Washington, where both taxa have been recorded. More detailed studies of the North American distributions of both these weeds are needed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cardamine constancei is known from Clearwater, Idaho, Kootenai, Nez Perce, and Shoshone counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 474. | FNA vol. 7, p. 471. |
Parent taxa | Brassicaceae > tribe Cardamineae > Cardamine | Brassicaceae > tribe Cardamineae > Cardamine |
Sibling taxa | ||
Synonyms | C. flexuosa subsp. debilis, C. flexuosa var. debilis, C. hirsuta subsp. flexuosa, C. scutata subsp. flexuosa | |
Name authority | Withering: Arr. Brit. Pl. ed. 3, 3: 578. (1796) | Detling: Madroño 3: 176, plate 9. (1935) |
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