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wavy bitter-cress, wood bitter-cress, woodland bitter-cress

Constance's bittercress

Habit Annuals or biennials; sparsely to densely hirsute basally or throughout, or glabrous. Perennials; glabrous or sparsely hirsute.
Rhizomes

absent.

cylindrical, slender, to 2 mm diam.

Stems

erect, ascending, or decumbent, branched or unbranched, (0.6–)1–5 dm, (slightly flexuous).

erect, unbranched, 1.5–5 dm, glabrous or sparsely hirsute basally.

Basal leaves

(often withered by anthesis), not rosulate, 5–15-foliolate, (2.7–)4–14(–19) cm, leaflets petiolulate;

petiole 0.7–5 cm, (ciliate or not);

lateral leaflet blade oblong, ovate, or elliptic, smaller than terminal, margins entire, repand, crenate, or 3 (or 5)-lobed;

terminal leaflet (petiolule 0.3–1.7 cm), blade reniform, broadly ovate, or suborbicular, 0.5–2.5 cm × 4–30 mm, margins repand, crenate, or 3 or 5-lobed.

Rhizomal leaves

absent.

Cauline leaves

3–15, 5–15-foliolate [leaves (2–)3.5–5.5(–7) cm, including petiole], petiolate, leaflets petiolulate;

petiole base not auriculate;

lateral leaflets similar to basal, (0.4–2.5 mm wide).

4–7, simple, (crowded distally), petiolate;

petiole (0.5–)1–4(–5) cm, base not auriculate;

blade usually broadly ovate or ovate-lanceolate to ovate-elliptic, rarely obovate, 5–13 cm × 20–65 mm, base cuneate, margins coarsely serrate (with apiculae), or repand to undulate (margins minutely pubescent, apex acute).

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, 1.5–2.5 × 0.7–1 mm, lateral pair not saccate basally;

petals white, spatulate, 2.5–4(–5) × 1–1.7 mm; (stamens rarely 4, lateral pair absent);

filaments 2–3 mm;

anthers ovate, 0.3–0.5 mm.

sepals oblong, 6–8 × 2–2.5 mm, lateral pair slightly saccate basally;

petals pink, oblanceolate, 15–28 × 5–8 mm, (claw to 10 mm, apex rounded);

filaments: median pairs 8–10 mm, lateral pair 4–6 mm;

anthers linear, 2–3 mm.

Fruiting pedicels

divaricate or ascending, (5–)6–14(–17) mm.

divaricate-ascending to suberect, 10–22 mm.

Fruits

linear, (torulose), (0.8–)1.2–2.8 cm × 1–1.5 mm;

ovules 18–40 per ovary;

style 0.3–1(–1.5) mm.

linear, 2.5–3.5(–5) cm × 1.9–2.1 mm;

ovules 12–16 per ovary;

style 2–3.5 mm.

Seeds

brown, oblong or subquadrate, 0.9–1.5 × 0.6–1 mm, (narrowly margined or not).

brown, oblong, 2–2.5 × 1.2–1.5 mm.

2n

= 32.

Cardamine flexuosa

Cardamine constancei

Phenology Flowering Apr–Jul. Flowering May–Jun.
Habitat Disturbed areas, fields, nurseries, plantations, gardens, flower beds, lawns, roadsides Moist cliffs, wooded creek bottoms, shaded draws, hillsides, moist woods, mixed coniferous forests, granitic soils
Elevation 0-1100 m (0-3600 ft) 400-600 m (1300-2000 ft)
Distribution
from FNA
AL; CA; FL; GA; IL; IN; LA; MD; MI; NC; NY; OH; OR; RI; TX; VA; WA; BC; NF; ON; Europe; e Asia [Introduced in North America; introduced also in Mexico, Central America, South America, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
ID
[WildflowerSearch map]
[BONAP county map]
Discussion

According to J. Lihová et al. (2006), the populations referred to Cardamine flexuosa in North America comprise two taxa of different polyploid origins and evolutionary histories: tetraploid C. flexuosa (2n = 32), native to Europe, and the octoploid taxon informally called “Asian C. flexuosa” (2n = 64), native to eastern Asia. For the latter, the name C. flexuosa subsp. debilis can be used. Nevertheless, these two taxa should be recognized at species level and the correct name for the Asian species should be sought. Based on available data, both taxa occupy the same habitats in North America, but the Asian taxon is much more widespread. The occurrence of European C. flexuosa was, until now, confirmed only for Washington, where both taxa have been recorded. More detailed studies of the North American distributions of both these weeds are needed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cardamine constancei is known from Clearwater, Idaho, Kootenai, Nez Perce, and Shoshone counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 474. FNA vol. 7, p. 471.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. flexuosa subsp. debilis, C. flexuosa var. debilis, C. hirsuta subsp. flexuosa, C. scutata subsp. flexuosa
Name authority Withering: Arr. Brit. Pl. ed. 3, 3: 578. (1796) Detling: Madroño 3: 176, plate 9. (1935)
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