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wavy bitter-cress, wood bitter-cress, woodland bitter-cress

bulb bittercress, bulbous bitter-cress, spring cress

Habit Annuals or biennials; sparsely to densely hirsute basally or throughout, or glabrous. Perennials; glabrous or sparsely pubescent distally.
Rhizomes

absent.

(tuberous at stem base, sometimes also at intervals) subglobose, lobed, 4–15 mm diam. (fleshy).

Stems

erect, ascending, or decumbent, branched or unbranched, (0.6–)1–5 dm, (slightly flexuous).

erect, unbranched, (1–)2–6 dm, glabrous or sparsely pubescent on distal 1/2 (trichomes 0.02–0.1 mm).

Basal leaves

(often withered by anthesis), not rosulate, 5–15-foliolate, (2.7–)4–14(–19) cm, leaflets petiolulate;

petiole 0.7–5 cm, (ciliate or not);

lateral leaflet blade oblong, ovate, or elliptic, smaller than terminal, margins entire, repand, crenate, or 3 (or 5)-lobed;

terminal leaflet (petiolule 0.3–1.7 cm), blade reniform, broadly ovate, or suborbicular, 0.5–2.5 cm × 4–30 mm, margins repand, crenate, or 3 or 5-lobed.

Rhizomal leaves

simple, (2–)4–13(–16) cm;

petiole (1.5–)2.5–10(–13) cm;

blade usually reniform to cordate or ovate, rarely oblong, (1–)2–4(–6) cm, base obtuse to cordate, margins usually repand or entire, rarely shallowly dentate.

Cauline leaves

3–15, 5–15-foliolate [leaves (2–)3.5–5.5(–7) cm, including petiole], petiolate, leaflets petiolulate;

petiole base not auriculate;

lateral leaflets similar to basal, (0.4–2.5 mm wide).

(2–)4–10(–14), simple, petiolate or sessile; (middle ones) shortly petiolate or (distally) sessile, base not auriculate;

blade ovate to oblong, or oblong-linear to lanceolate, 3–6(–9) cm × 10–30(–45) mm, margins entire, repand, or dentate (margins minutely pubescent).

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, 1.5–2.5 × 0.7–1 mm, lateral pair not saccate basally;

petals white, spatulate, 2.5–4(–5) × 1–1.7 mm; (stamens rarely 4, lateral pair absent);

filaments 2–3 mm;

anthers ovate, 0.3–0.5 mm.

sepals oblong, 2.5–4.5 × 1.5–2 mm, lateral pair not saccate basally, (glabrous);

petals usually white, rarely pale pink, obovate, (6–)7–12(–16) × 3–5 mm, (short-clawed, apex rounded);

filaments: median pairs 4.5–7 mm, lateral pair 2–3.5 mm;

anthers oblong, 1–1.5 mm.

Fruiting pedicels

divaricate or ascending, (5–)6–14(–17) mm.

ascending to divaricate, (10–)15–22(–30) mm.

Fruits

linear, (torulose), (0.8–)1.2–2.8 cm × 1–1.5 mm;

ovules 18–40 per ovary;

style 0.3–1(–1.5) mm.

linear, 2–3.5(–4) cm × 1.4–1.7 mm;

ovules 14–24 per ovary;

style 2–4(–5) mm.

Seeds

brown, oblong or subquadrate, 0.9–1.5 × 0.6–1 mm, (narrowly margined or not).

dark orange to greenish yellow, oblong or globose, 1.7–2.1 × 1–1.4 mm.

2n

= 32.

= 16, 56, 64, 80, 96, 112.

Cardamine flexuosa

Cardamine bulbosa

Phenology Flowering Apr–Jul. Flowering Mar–Jun.
Habitat Disturbed areas, fields, nurseries, plantations, gardens, flower beds, lawns, roadsides Wet grounds, low woodland, moss hummocks, alluvial woods, grassy floodplains, wet pastures, meadows, pinelands, creek bottoms, stream banks, sandy bottoms, ditches, mesic or wet forests, swamps, marshes, seepy bluffs
Elevation 0-1100 m (0-3600 ft) 0-900 m (0-3000 ft)
Distribution
from FNA
AL; CA; FL; GA; IL; IN; LA; MD; MI; NC; NY; OH; OR; RI; TX; VA; WA; BC; NF; ON; Europe; e Asia [Introduced in North America; introduced also in Mexico, Central America, South America, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; SD; TN; TX; VA; VT; WI; WV; MB; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

According to J. Lihová et al. (2006), the populations referred to Cardamine flexuosa in North America comprise two taxa of different polyploid origins and evolutionary histories: tetraploid C. flexuosa (2n = 32), native to Europe, and the octoploid taxon informally called “Asian C. flexuosa” (2n = 64), native to eastern Asia. For the latter, the name C. flexuosa subsp. debilis can be used. Nevertheless, these two taxa should be recognized at species level and the correct name for the Asian species should be sought. Based on available data, both taxa occupy the same habitats in North America, but the Asian taxon is much more widespread. The occurrence of European C. flexuosa was, until now, confirmed only for Washington, where both taxa have been recorded. More detailed studies of the North American distributions of both these weeds are needed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 474. FNA vol. 7, p. 469.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. flexuosa subsp. debilis, C. flexuosa var. debilis, C. hirsuta subsp. flexuosa, C. scutata subsp. flexuosa Arabis bulbosa, Arabis rhomboidea, C. rhomboidea, C. rhomboidea var. hirsuta, C. rhomboidea var. parviflora, C. rhomboidea var. pilosa, Dentaria rhomboidea, Dracamine bulbosa
Name authority Withering: Arr. Brit. Pl. ed. 3, 3: 578. (1796) (Schreber ex Muhlenberg) Britton: Prelim. Cat., 4. (1888)
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