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limestone bittercress, pink bitter-cress

cliff bittercress

Habit Perennials; hirsute throughout or glabrous proximally. Perennials; glabrous throughout.
Rhizomes

(tuberous at stem base), subglobose, (lobed or not), (3–)4–10 mm diam., (fleshy).

cylindrical, slender, 1–2 mm diam.

Stems

erect, unbranched, (0.7–)1–2.5(–3) dm, sparsely to densely hirsute, or glabrous basally, (trichomes (0.2–)0.3–0.6(–0.8) mm).

erect or decumbent at base, unbranched, 0.6–2 dm.

Rhizomal leaves

simple, (3–)5–15(–18) cm;

petiole (2–)4–12(–16) cm;

blade often orbicular to cordate, sometimes reniform or ovate, (1–)2–6 cm × (7–)17–50 mm, base obtuse to cordate, margins repand or entire.

palmately or subpalmately compound, 3 or 5 (or 7)-foliolate, 5.5–17(–22) cm, (fleshy), petiolate, leaflets petiolulate or subsessile;

petiole 4–14(–17) cm;

lateral leaflets subsessile, blade similar to terminal, sometimes smaller;

terminal leaflet (petiolule 0.1–0.5 cm), blade ovate to lanceolate or elliptic-oblong, 1–3 cm × 6–17 mm, base cuneate or obtuse, margins entire, (apiculate).

Cauline leaves

3–6(–8), simple, petiolate or sessile; (middle) shortly petiolate or (distal) sessile, base not auriculate;

blade oblong to ovate or lanceolate, 2–5 cm × 5–25 mm, margins entire, repand, or coarsely dentate.

2 or 3, 3 or 5-foliolate, petiolate, leaflets petiolulate or sessile;

petiole 0.7–4(–8) cm, base not auriculate;

lateral leaflets sessile, blade similar to terminal, smaller;

terminal leaflet petiolulate (0.1–0.5 cm), blade elliptic to oblong, or ovate, 1.2–3.5 cm × 4–25 mm, margins entire.

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, 2.5–4(–6) × 1.5–2.5 mm, lateral pair not saccate basally, (surfaces often hirsute);

petals usually rose-purple to pink, rarely white, obovate, (7–)8–13(–15) × 3–5 mm, (short-clawed, apex rounded);

filaments: median pairs 4–7 mm, lateral pair 2–4 mm;

anthers oblong, 1.3–1.7 mm.

sepals oblong, 3–5 × 1.5–2 mm, lateral pair saccate basally;

petals white, obovate, 8–14 × 4–7 mm, (short-clawed, apex rounded or subemarginate);

filaments: median pairs 4–5 mm, lateral pair 2.5–3.5 mm;

anthers oblong, 1–1.2 mm.

Fruiting pedicels

ascending to divaricate, (10–)15–35(–50) mm, sparsely pubescent or glabrous.

ascending to divaricate, 6–18 mm.

Fruits

linear, (1.5–)2–4 cm × 1.5–2 mm;

ovules 10–16 per ovary;

style 2–5 mm.

linear, 2–4 cm × 1.5–2.2 mm;

ovules 10–14 per ovary;

style 1–5 mm.

Seeds

brown, oblong to ovoid, 1.7–2.5 × 1–5 mm.

brown, oblong, 1.8–2.2 × 1.2–1.5 mm.

2n

= 56, 64, 96, 112, 144.

Cardamine douglassii

Cardamine rupicola

Phenology Flowering Mar–May. Flowering Jul–Aug.
Habitat Rich woods, bluffs, mesic bottomland forests, rocky hillsides, floodplains, seepage of bogs, springy areas Limestone talus slopes, loose limey shale, moist banks
Elevation 50-400 m (200-1300 ft) 2200-2700 m (7200-8900 ft)
Distribution
from FNA
AL; CT; DC; IA; IL; IN; KY; MA; MD; MI; MN; MO; NC; NJ; NY; OH; PA; SC; TN; VA; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
MT
[BONAP county map]
Discussion

Cardamine rupicola is known from Flathead, Lewis and Clarke, and Missoula counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 473. FNA vol. 7, p. 483.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. umbellata
Synonyms Arabis rhomboidea var. purpurea, Dentaria douglassii, Dracamine purpurea, Thlaspi tuberosum C. californica var. rupicola, Dentaria rupicola
Name authority Britton: Trans. New York Acad. Sci. 9: 8. (1889) (O. E. Schulz) C. L. Hitchcock: in C. L. Hitchcock et al., Vasc. Pl. Pacif. N.W. 2: 474. (1964)
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