Cardamine douglassii |
Cardamine macrocarpa |
|
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limestone bittercress, pink bitter-cress |
largeseed bittercress |
|
Habit | Perennials; hirsute throughout or glabrous proximally. | Annuals; glabrous or sparsely puberulent. |
Rhizomes | (tuberous at stem base), subglobose, (lobed or not), (3–)4–10 mm diam., (fleshy). |
absent. |
Stems | erect, unbranched, (0.7–)1–2.5(–3) dm, sparsely to densely hirsute, or glabrous basally, (trichomes (0.2–)0.3–0.6(–0.8) mm). |
(simple or several from base), erect or decumbent, (flexuous or straight, narrowly winged-angled), unbranched or branched distally, (1.4–)2–4.5(–5.3) dm. |
Basal leaves | (soon withered), not rosulate. |
|
Rhizomal leaves | simple, (3–)5–15(–18) cm; petiole (2–)4–12(–16) cm; blade often orbicular to cordate, sometimes reniform or ovate, (1–)2–6 cm × (7–)17–50 mm, base obtuse to cordate, margins repand or entire. |
|
Cauline leaves | 3–6(–8), simple, petiolate or sessile; (middle) shortly petiolate or (distal) sessile, base not auriculate; blade oblong to ovate or lanceolate, 2–5 cm × 5–25 mm, margins entire, repand, or coarsely dentate. |
3–9, middle ones 5–9-foliolate, petiolate, leaflets petiolulate or subsessile; petiole 1–3 cm, base not auriculate; lateral leaflets similar to terminal, blade often smaller, with oblique base, distal leaflets subsessile, blade smaller and narrower distally; terminal leaflet (petiolule 0.2–0.8 cm), blade usually broadly ovate to narrowly lanceolate, rarely oblong, 0.7–2 cm × 2–10 mm, base cuneate to rounded, margins repand, crenate, or 3-lobed. |
Racemes | ebracteate. |
ebracteate, (rachis slightly to strongly flexuous or geniculate). |
Flowers | sepals oblong, 2.5–4(–6) × 1.5–2.5 mm, lateral pair not saccate basally, (surfaces often hirsute); petals usually rose-purple to pink, rarely white, obovate, (7–)8–13(–15) × 3–5 mm, (short-clawed, apex rounded); filaments: median pairs 4–7 mm, lateral pair 2–4 mm; anthers oblong, 1.3–1.7 mm. |
sepals oblong, 2–3.5 × 0.7–1.2 mm, lateral pair not saccate basally; petals white, linear, 6–8 × 0.7–1 mm; filaments: median pairs 4–5 mm, lateral pair 3.5–4 mm; anthers oblong, 0.7–1 mm. |
Fruiting pedicels | ascending to divaricate, (10–)15–35(–50) mm, sparsely pubescent or glabrous. |
horizontal to divaricate or ascending, (3–)4–9(–12) mm. |
Fruits | linear, (1.5–)2–4 cm × 1.5–2 mm; ovules 10–16 per ovary; style 2–5 mm. |
linear, (2.5–)3–4.6 cm × 1.7–2.1 mm; ovules 14–22 per ovary; style 1–3 mm. |
Seeds | brown, oblong to ovoid, 1.7–2.5 × 1–5 mm. |
dark brown, oblong, 2–2.5 × 0.9–1.2 mm. |
2n | = 56, 64, 96, 112, 144. |
|
Cardamine douglassii |
Cardamine macrocarpa |
|
Phenology | Flowering Mar–May. | Flowering Mar–Sep. |
Habitat | Rich woods, bluffs, mesic bottomland forests, rocky hillsides, floodplains, seepage of bogs, springy areas | Rock crevices and ledges, gravel bars of mountain streams, moist rocky stream banks, shaded loamy forest floors |
Elevation | 50-400 m (200-1300 ft) | |
Distribution |
AL; CT; DC; IA; IL; IN; KY; MA; MD; MI; MN; MO; NC; NJ; NY; OH; PA; SC; TN; VA; WI; WV; ON
|
TX; Mexico (Coahuila, Nuevo León) |
Discussion | Cardamine macrocarpa is known from the Chisos Mountains, Brewster County. The characters by which var. texana is said to differ from var. macrocarpa are artificially drawn, and the style length, presence or absence of indumentum on the pedicels, and degree of flexuosity of the raceme rachises do not correlate and can vary within a given area. For these reasons, we do not recognize infraspecific taxa in Cardamine macrocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 473. | FNA vol. 7, p. 476. |
Parent taxa | Brassicaceae > tribe Cardamineae > Cardamine | Brassicaceae > tribe Cardamineae > Cardamine |
Sibling taxa | ||
Synonyms | Arabis rhomboidea var. purpurea, Dentaria douglassii, Dracamine purpurea, Thlaspi tuberosum | C. macrocarpa var. texana |
Name authority | Britton: Trans. New York Acad. Sci. 9: 8. (1889) | Brandegee: Zoë 5: 233. (1906) |
Web links |