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limestone bittercress, pink bitter-cress

long's bitter-cress

Habit Perennials; hirsute throughout or glabrous proximally. Perennials; glabrous throughout.
Rhizomes

(tuberous at stem base), subglobose, (lobed or not), (3–)4–10 mm diam., (fleshy).

cylindrical, slender, 0.8–1.5 mm diam.

Stems

erect, unbranched, (0.7–)1–2.5(–3) dm, sparsely to densely hirsute, or glabrous basally, (trichomes (0.2–)0.3–0.6(–0.8) mm).

prostrate to decumbent or erect, unbranched or branched, 0.5–4(–6) dm.

Rhizomal leaves

simple, (3–)5–15(–18) cm;

petiole (2–)4–12(–16) cm;

blade often orbicular to cordate, sometimes reniform or ovate, (1–)2–6 cm × (7–)17–50 mm, base obtuse to cordate, margins repand or entire.

absent.

Cauline leaves

3–6(–8), simple, petiolate or sessile; (middle) shortly petiolate or (distal) sessile, base not auriculate;

blade oblong to ovate or lanceolate, 2–5 cm × 5–25 mm, margins entire, repand, or coarsely dentate.

3–10, usually simple, rarely 3 or 5-foliolate, petiolate; petiolulate or sessile;

petiole 0.4–2.5 cm, base not auriculate;

lateral leaflets (when present) sessile or petiolulate (to 0.2 cm), blade similar to terminal, considerably smaller;

terminal leaflet blade orbicular to reniform or ovate to oblong, 0.4–3 cm × 3–20 mm, (somewhat fleshy), base subcordate to rounded or subtruncate, margins entire or repand, rarely undulate, (apex rounded).

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, 2.5–4(–6) × 1.5–2.5 mm, lateral pair not saccate basally, (surfaces often hirsute);

petals usually rose-purple to pink, rarely white, obovate, (7–)8–13(–15) × 3–5 mm, (short-clawed, apex rounded);

filaments: median pairs 4–7 mm, lateral pair 2–4 mm;

anthers oblong, 1.3–1.7 mm.

sepals ovate, 0.7–1.2 × 0.5–0.7 mm, lateral pair not saccate basally;

petals absent or rudimentary, to 0.7 mm;

filaments 0.5–0.8 mm;

anthers ovate, 0.1–0.2 mm.

Fruiting pedicels

ascending to divaricate, (10–)15–35(–50) mm, sparsely pubescent or glabrous.

divaricate-ascending to spreading, 0.5–2(–4) mm.

Fruits

linear, (1.5–)2–4 cm × 1.5–2 mm;

ovules 10–16 per ovary;

style 2–5 mm.

narrowly oblong to linear, (3–)5–10(–15) × 0.8–1.2 mm;

ovules 6–22 per ovary;

style 0.2–0.5(–1) mm.

Seeds

brown, oblong to ovoid, 1.7–2.5 × 1–5 mm.

light or yellowish brown, oblong or ovoid, 1–1.4 × 0.7–1 mm.

2n

= 56, 64, 96, 112, 144.

Cardamine douglassii

Cardamine longii

Phenology Flowering Mar–May. Flowering Jun–Sep.
Habitat Rich woods, bluffs, mesic bottomland forests, rocky hillsides, floodplains, seepage of bogs, springy areas Tidal marshes, mud flats, tidal shores of rivers, shallow water, swampy areas, shady rocky crevices covered at high tide
Elevation 50-400 m (200-1300 ft) 0-10 m (0-0 ft)
Distribution
from FNA
AL; CT; DC; IA; IL; IN; KY; MA; MD; MI; MN; MO; NC; NJ; NY; OH; PA; SC; TN; VA; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; MA; MD; ME; NC; NJ; NY; SC; VA
[BONAP county map]
Source FNA vol. 7, p. 473. FNA vol. 7, p. 476.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms Arabis rhomboidea var. purpurea, Dentaria douglassii, Dracamine purpurea, Thlaspi tuberosum
Name authority Britton: Trans. New York Acad. Sci. 9: 8. (1889) Fernald: Rhodora 19: 91. (1917)
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