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Constance's bittercress

Holmgren's bitter-cress

Habit Perennials; glabrous or sparsely hirsute. Perennials; glabrous throughout.
Rhizomes

cylindrical, slender, to 2 mm diam.

slender, 0.5–1 mm diam.

Stems

erect, unbranched, 1.5–5 dm, glabrous or sparsely hirsute basally.

erect to ascending, branched, 1.5–2 dm.

Rhizomal leaves

absent.

Cauline leaves

4–7, simple, (crowded distally), petiolate;

petiole (0.5–)1–4(–5) cm, base not auriculate;

blade usually broadly ovate or ovate-lanceolate to ovate-elliptic, rarely obovate, 5–13 cm × 20–65 mm, base cuneate, margins coarsely serrate (with apiculae), or repand to undulate (margins minutely pubescent, apex acute).

3 or 4, 3- or 5-foliolate, petiolate, leaflets petiolulate;

petiole 0.5–1.7 cm, base not auriculate;

lateral leaflet blade linear to linear-lanceolate, 0.9–1.5 cm × 1–3 mm, margins entire;

terminal leaflet (petiolule 0.3–0.5 cm), blade obovate to lanceolate (linear distally), 1.3–2.3 cm × 2–8 mm, base cuneate, margins entire or subapically 1- or 2-toothed.

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, 6–8 × 2–2.5 mm, lateral pair slightly saccate basally;

petals pink, oblanceolate, 15–28 × 5–8 mm, (claw to 10 mm, apex rounded);

filaments: median pairs 8–10 mm, lateral pair 4–6 mm;

anthers linear, 2–3 mm.

sepals oblong, 1.2–1.5 × 0.6–0.8 mm, lateral pair not saccate basally;

petals white, oblanceolate, 2–2.5 × 0.8–1 mm, (not clawed, apex rounded);

filaments: median pairs 1.5–1.8 mm, lateral pair 1–1.2 mm;

anthers ovate, 0.2–0.3 mm.

Fruiting pedicels

divaricate-ascending to suberect, 10–22 mm.

ascending to suberect, 5–14 mm.

Fruits

linear, 2.5–3.5(–5) cm × 1.9–2.1 mm;

ovules 12–16 per ovary;

style 2–3.5 mm.

linear, 1–2 cm × ca. 1 mm;

ovules 16–24 per ovary;

style 0.5–0.7 mm.

Seeds

brown, oblong, 2–2.5 × 1.2–1.5 mm.

brown, oblong, 1.2–1.5 × 0.8–1 mm.

Rhizomal

and basal leaves absent.

Cardamine constancei

Cardamine holmgrenii

Phenology Flowering May–Jun. Flowering Jul–Aug.
Habitat Moist cliffs, wooded creek bottoms, shaded draws, hillsides, moist woods, mixed coniferous forests, granitic soils Boggy slopes
Elevation 400-600 m (1300-2000 ft)
Distribution
from FNA
ID
[WildflowerSearch map]
[BONAP county map]
from FNA
OR
[BONAP county map]
Discussion

Cardamine constancei is known from Clearwater, Idaho, Kootenai, Nez Perce, and Shoshone counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cardamine holmgrenii is known only from the type collection, from the Blue Mountains in Baker County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 471. FNA vol. 7, p. 475.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Name authority Detling: Madroño 3: 176, plate 9. (1935) Al-Shehbaz: Harvard Pap. Bot. 11: 275. (2007)
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