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Constance's bittercress

heart-leaf bitter-cress, large mountain bitter-cress, Lyall's bitter-cress

Habit Perennials; glabrous or sparsely hirsute. Perennials; glabrous or densely puberulent.
Rhizomes

cylindrical, slender, to 2 mm diam.

cylindrical, slender or stout, 1.5–3 mm diam., (not fleshy).

Stems

erect, unbranched, 1.5–5 dm, glabrous or sparsely hirsute basally.

erect, unbranched or branched distally, 2–7(–10.2) dm, glabrous or sparsely to densely puberulent basally or throughout.

Rhizomal leaves

absent.

usually absent, rarely present, simple, 5–15 cm;

petiole 2.5–12 cm;

blade reniform to cordate, 1.5–4.5(–6) cm.

Cauline leaves

4–7, simple, (crowded distally), petiolate;

petiole (0.5–)1–4(–5) cm, base not auriculate;

blade usually broadly ovate or ovate-lanceolate to ovate-elliptic, rarely obovate, 5–13 cm × 20–65 mm, base cuneate, margins coarsely serrate (with apiculae), or repand to undulate (margins minutely pubescent, apex acute).

(3–)5–17(–23), simple or, rarely, proximalmost 3-foliolate, petiolate;

petiole 1–6.5(–8.5) cm, base not auriculate;

blade reniform, deltate-cordate, or ovate-cordate, (1–)2–7.2(–9.7) cm × 10–55(–85) mm, (somewhat fleshy), base cordate or truncate, margins crenate to slightly sinuate, (veins ending in apiculae, surfaces glabrous or sparsely to densely pubescent).

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, 6–8 × 2–2.5 mm, lateral pair slightly saccate basally;

petals pink, oblanceolate, 15–28 × 5–8 mm, (claw to 10 mm, apex rounded);

filaments: median pairs 8–10 mm, lateral pair 4–6 mm;

anthers linear, 2–3 mm.

sepals oblong, 2.5–4.5 × 1.5–2 mm, lateral pair slightly saccate basally;

petals white, broadly obovate, 7–12 × 4–6 mm (claw to 6 mm, apex rounded to subtruncate or emarginate);

filaments: median pairs 3.5–5 mm, lateral pair 2–3.5 mm;

anthers oblong, 1–1.5 mm.

Fruiting pedicels

divaricate-ascending to suberect, 10–22 mm.

divaricate to ascending, (7–)10–20 mm.

Fruits

linear, 2.5–3.5(–5) cm × 1.9–2.1 mm;

ovules 12–16 per ovary;

style 2–3.5 mm.

linear, (2–)2.5–3.7(–4) cm × 1.2–2 mm; (valves glabrous or sparsely puberulent);

ovules 14–24 per ovary;

style 0.5–3(–6) mm.

Seeds

brown, oblong, 2–2.5 × 1.2–1.5 mm.

brown, oblong, 1.6–2 × 1–1.3 mm.

2n

= 24.

Cardamine constancei

Cardamine cordifolia

Phenology Flowering May–Jun. Flowering May–Aug.
Habitat Moist cliffs, wooded creek bottoms, shaded draws, hillsides, moist woods, mixed coniferous forests, granitic soils Stream banks, springs, shady gullies, creek bottoms, lakeshores, ponds, cold springs, meadows, moist hillsides, mossy areas, alpine streams, mixed coniferous forests
Elevation 400-600 m (1300-2000 ft) 600-3600 m (2000-11800 ft)
Distribution
from FNA
ID
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[BONAP county map]
from FNA
AZ; CA; CO; ID; NM; NV; OR; UT; WA; WY; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Cardamine constancei is known from Clearwater, Idaho, Kootenai, Nez Perce, and Shoshone counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cardamine cordifolia is highly variable in leaf morphology, especially in leaf width, depth of the cordate base, and indumentum. This variation occurs throughout the species range and is rather weakly or not at all correlated with geography. In the absence of a detailed biosystematic study over the entire species range, we follow N. H. Holmgren (2005b) in not recognizing any infraspecific taxa, instead of accepting the three rather poorly defined varieties recognized by R. C. Rollins (1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 471. FNA vol. 7, p. 471.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms C. cordifolia var. cardiophylla, C. cordifolia var. diversifolia, C. cordifolia var. incana, C. cordifolia subsp. lyallii, C. cordifolia var. lyallii, C. cordifolia var. pubescens, C. incana, C. infausta, C. lyallii, C. uintahensis
Name authority Detling: Madroño 3: 176, plate 9. (1935) A. Gray: Mem. Amer. Acad. Arts, n. s., 4: 8. (1849)
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