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cut-leaf toothwort

alpine bitter-cress, few-seed bitter-cress, Idaho bittercress, little western bitter-cress, Siberian bitter-cress

Habit Perennials; usually sparsely pubescent, sometimes glabrous. Annuals or biennials; usually sparsely hirsute (at least proximally), rarely glabrous.
Rhizomes

(tuberiform, fragile), moniliform, segments fusiform, 2–10(–20) mm diam. (fleshy).

absent.

Stems

erect, unbranched, (1–)2–4(–5.5) dm, glabrous or pubescent distally.

(simple or few from base), erect to ascending, (not flexuous), unbranched or branched distally, (0.5–)0.8–3.2(–4.1) dm, usually pubescent throughout or proximally, rarely glabrous.

Basal leaves

(persistent to anthesis), rosulate, pinnately compound, 5–9(–13)-foliolate, 2–8.5(–11) cm, leaflets petiolulate or subsessile;

petiole 1–6 cm;

lateral leaflets petiolulate or subsessile, blade obovate to oblanceolate, smaller than terminal, margins entire or crenate;

terminal leaflet (petiolule 0.1–0.7 cm), blade usually orbicular to ovate, rarely oblong, 0.4–1.5(–2.3) cm × 3–10(–13) mm, base cordate to rounded, margins entire, or crenate-dentate to obscurely 3 or 5-lobed, (surfaces often hirsute, sometimes glabrous).

Rhizomal leaves

3-foliolate, (7–)10–20(–30) cm, leaflets sessile;

petiole (4–)7–18(–25) cm;

lateral leaflets similar to terminal, blade sometimes smaller;

terminal leaflet blade oblong, lanceolate, oblanceolate, or linear, 2.5–6 cm, base cuneate, margins coarsely dentate to incised, laciniate, or 3-lobed (lobes usually toothed to incised, rarely entire, surfaces puberulent or not, trichomes 0.2–0.3 mm).

Cauline leaves

(2 or) 3, 3-foliolate (usually whorled or opposite, rarely alternate, similar in morphology to rhizomal leaves), petiolate, leaflets petiolulate or subsessile;

petiole (1–)1.5–6(–7.5) cm, base not auriculate;

lateral leaflets sessile, blade similar to terminal, sometimes smaller;

terminal leaflet subsessile or petiolulate (to 3 cm), blade lanceolate, linear, or oblanceolate, (3–)4–10(–12) cm × (3–)5–20(–25) mm, margins usually coarsely dentate to incised, rarely subentire (margins minutely puberulent).

3–8, pinnately compound, similar to basal, smaller and fewer distally, petiolate, leaflets petiolulate or subsessile;

petiole 0.5–2 cm, base not auriculate.

Racemes

ebracteate.

ebracteate.

Flowers

sepals oblong, (4–)5–8 × 2–4 mm, lateral pair slightly saccate basally;

petals white to pale pink, oblanceolate, (8–)10–20 × (3–)4–7(–9) mm, (short-clawed, apex rounded);

filaments: median pairs 8–12 mm, lateral pair 6–8 mm;

anthers oblong-linear, 1.5–2.5 mm.

sepals oblong, 1.3–1.8(–2) × 0.5–1 mm, lateral pair not saccate basally;

petals white, narrowly spatulate to oblanceolate, 2.5–3.5 × 0.9–1.5 mm (not clawed);

filaments: median pairs 1.7–2.5 mm, lateral pair 1.2–2 mm;

anthers ovate, 0.3–0.5 mm.

Fruiting pedicels

ascending to divaricate, (6–)10–27(–33) mm.

divaricate-ascending, (2–)3–9(–12) mm.

Fruits

linear-lanceolate, (2–)2.5–3.8(–4.8) cm × 1.5–3 mm; (valves glabrous or sparsely pubescent);

ovules 10–14 per ovary;

style (2–)5–9(–12) mm.

linear, (torulose), (1.3–)1.6–2.8 cm × 1–1.7 mm; (valves glabrous or sparsely pubescent);

ovules 16–36(–42) per ovary;

style 0.4–1(–1.5) mm.

Seeds

brown, oblong, 1.6–3 × 1.8–2 mm.

brown, oblong, 1–1.6 × 0.8–1.2 mm.

2n

= 128–256.

= 16.

Cardamine concatenata

Cardamine oligosperma

Phenology Flowering Feb–May. Flowering Mar–Jul.
Habitat Wooded bottoms and bluffs, rich woods, limestone cliffs and outcrops, rocky banks, mesic forests, moist areas with leaf litter, floodplain woods Stream banks, shady banks, creek bottoms, lakeshores, meadows, moist areas, wooded slopes
Elevation 0-1000 m (0-3300 ft) 50-3300 m (200-10800 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; VT; WI; WV; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY; BC; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 7, p. 470. FNA vol. 7, p. 479.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. diphylla, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms Dentaria concatenata, C. laciniata var. integra, C. laciniata var. lasiocarpa, Dentaria laciniata, Dentaria laciniata var. alterna, Dentaria laciniata var. coalescens, Dentaria laciniata var. integra, Dentaria laciniata var. latifolia, Dentaria laciniata var. opposita C. acuminata, C. hirsuta var. acuminata, C. hirsuta var. bracteata, C. hirsuta subsp. oligosperma, C. hirsuta var. parviflora, C. oligosperma var. bracteata, C. oligosperma var. lucens, C. oligosperma var. unijuga, C. unijuga
Name authority (Michaux) O. Schwarz: Repert. Spec. Nov. Regni Veg. 46: 188. (1939) Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 85. (1838)
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