Cardamine concatenata |
Cardamine maxima |
|
---|---|---|
cut-leaf toothwort |
large toothwort |
|
Habit | Perennials; usually sparsely pubescent, sometimes glabrous. | Perennials; glabrous (except leaflet margins and, sometimes, rachis). |
Rhizomes | (tuberiform, fragile), moniliform, segments fusiform, 2–10(–20) mm diam. (fleshy). |
cylindrical, 3–6 mm diam., (distinctly constricted at intervals, non-uniform diam., fleshy, slightly fragile, with dentate leaf scars). |
Stems | erect, unbranched, (1–)2–4(–5.5) dm, glabrous or pubescent distally. |
erect, unbranched, 0.9–3(–4) dm. |
Rhizomal leaves | 3-foliolate, (7–)10–20(–30) cm, leaflets sessile; petiole (4–)7–18(–25) cm; lateral leaflets similar to terminal, blade sometimes smaller; terminal leaflet blade oblong, lanceolate, oblanceolate, or linear, 2.5–6 cm, base cuneate, margins coarsely dentate to incised, laciniate, or 3-lobed (lobes usually toothed to incised, rarely entire, surfaces puberulent or not, trichomes 0.2–0.3 mm). |
3-foliolate, 7–20 cm, leaflets petiolulate or subsessile; petiole 4–15 cm; lateral leaflets subsessile or petiolulate, blade often similar to terminal, base often oblique; terminal leaflet (petiolule 0.2–1(–1.7) cm), blade broadly ovate to oblong, 2–7.5 cm × 12–37 mm, base cuneate to obtuse, margins coarsely dentate to sharply incised, or deeply cleft into 2 or 3 lobes (lobes dentate or incised, margins puberulent). |
Cauline leaves | (2 or) 3, 3-foliolate (usually whorled or opposite, rarely alternate, similar in morphology to rhizomal leaves), petiolate, leaflets petiolulate or subsessile; petiole (1–)1.5–6(–7.5) cm, base not auriculate; lateral leaflets sessile, blade similar to terminal, sometimes smaller; terminal leaflet subsessile or petiolulate (to 3 cm), blade lanceolate, linear, or oblanceolate, (3–)4–10(–12) cm × (3–)5–20(–25) mm, margins usually coarsely dentate to incised, rarely subentire (margins minutely puberulent). |
2 or 3, 3-foliolate, (rarely subopposite), petiolate, leaflets petiolulate or subsessile; petiole (0.5–)1–4(–6.5) cm, base not auriculate; lateral and terminal leaflets similar to rhizomal, distalmost sometimes much smaller. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, (4–)5–8 × 2–4 mm, lateral pair slightly saccate basally; petals white to pale pink, oblanceolate, (8–)10–20 × (3–)4–7(–9) mm, (short-clawed, apex rounded); filaments: median pairs 8–12 mm, lateral pair 6–8 mm; anthers oblong-linear, 1.5–2.5 mm. |
sepals (erect to ascending), oblong, 5–7 × 2–3 mm, lateral pair slightly saccate basally; petals white or pink, oblanceolate, 10–17 × 3–6 mm, (not clawed, apex rounded); filaments: median pairs 4–8 mm, lateral pair 3–6.5 mm; anthers linear, 1.7–2.7 mm. |
Fruiting pedicels | ascending to divaricate, (6–)10–27(–33) mm. |
(flowering ones) horizontal to divaricate or deflexed, 7–20 mm. |
Fruits | linear-lanceolate, (2–)2.5–3.8(–4.8) cm × 1.5–3 mm; (valves glabrous or sparsely pubescent); ovules 10–14 per ovary; style (2–)5–9(–12) mm. |
(undeveloped), linear-lanceolate, to 3 cm × 2 mm; ovules 10–14 per ovary; style 3.5–7 mm. |
Seeds | brown, oblong, 1.6–3 × 1.8–2 mm. |
not known. |
2n | = 128–256. |
= 120, 124, 132, 138, 156, 161, ca. 208. |
Cardamine concatenata |
Cardamine maxima |
|
Phenology | Flowering Feb–May. | Flowering Apr–Jun. |
Habitat | Wooded bottoms and bluffs, rich woods, limestone cliffs and outcrops, rocky banks, mesic forests, moist areas with leaf litter, floodplain woods | Rich woods, shady ravines, ledges, moist alluvial bottoms, steep forested slopes, stream banks |
Elevation | 0-1000 m (0-3300 ft) | |
Distribution |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; VT; WI; WV; ON; QC
|
CT; MA; ME; MI; NJ; NY; OH; PA; VT; NB; ON; QC
|
Discussion | Cardamine maxima has not been found with mature fruits and seeds and it has long been suspected to be a hybrid between C. concatenata and C. diphylla. Molecular studies (P. W. Sweeney and R. A. Price 2000) indicate that C. maxima is distinct from both those species. Although we hesitate to maintain it as a species, its wide distribution and morphological distinctness warrant its recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 470. | FNA vol. 7, p. 477. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Dentaria concatenata, C. laciniata var. integra, C. laciniata var. lasiocarpa, Dentaria laciniata, Dentaria laciniata var. alterna, Dentaria laciniata var. coalescens, Dentaria laciniata var. integra, Dentaria laciniata var. latifolia, Dentaria laciniata var. opposita | Dentaria maxima, C. anomala, Dentaria anomala |
Name authority | (Michaux) O. Schwarz: Repert. Spec. Nov. Regni Veg. 46: 188. (1939) | (Nuttall) Alph. Wood: Amer. Bot. Fl., 38. (1870) |
Web links |