Cardamine clematitis |
Cardamine angustata |
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small mountain bittercress |
slender toothwort |
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Habit | Perennials; glabrous throughout. | Perennials; usually glabrous, rarely sparsely pubescent. |
Rhizomes | cylindrical, slender, 1–3 mm diam. |
(tuberiform, fragile) moniliform, segments fusiform, 3–6 mm diam. (fleshy). |
Stems | erect, unbranched or rarely branched distally, (0.8–)1–2.5(–3.5) dm. |
erect, unbranched, 1.2–3(–4) dm, glabrous or pubescent. |
Rhizomal leaves | simple or 3-foliolate, (1.5–)3–8 cm, leaflets petiolulate or subsessile; petiole (1–)2–6 cm; lateral leaflets subsessile, blade similar to terminal, much smaller; terminal leaflet petiolulate (to 1 cm), blade reniform to cordate, (0.5–)1–2 cm, base cordate, margins entire or 3-lobed. |
3-foliolate, to 24 cm, leaflets petiolulate or subsessile; petiole (3–)5–12(–16) cm; lateral leaflets subsessile or petiolulate (0.2–1 cm), blade similar to terminal leaflet or smaller; terminal leaflet (petiolule) (0.2–)0.5–1.5(–2) cm, blade broadly ovate to rhombic-obovate, 1.5–6(–8) cm, base usually cuneate, rarely subtruncate, margins coarsely dentate to crenate or 3-lobed, surfaces puberulent or not. |
Cauline leaves | 3–7, 3-foliolate (or distalmost simple), petiolate, leaflets petiolulate or sessile; petiole 0.7–3.5 cm, base auriculate (auricle 0.7–5 mm); lateral leaflets sessile or petiolulate (to 0.5 cm), blade oblong to ovate or oblong; terminal leaflet (petiolule 0.5–1 cm), blade broadly ovate to suborbicular or reniform, 1.5–4 cm × 15–35 mm, margins often 3 or 5-lobed (lobe apex minutely apiculate, glabrous on margin). |
2 (or 3), 3-foliolate (usually alternate, rarely opposite, different in morphology from rhizomal), petiolate, leaflets petiolulate or sessile; petiole 0.5–2 cm, base not auriculate; lateral leaflets sessile, blade similar to terminal, smaller, margins usually dentate, rarely entire; terminal leaflet sessile or petiolulate, blade narrowly lanceolate to narrowly oblong or oblong-lanceolate, 2–7 × 0.3–0.6 cm, margins minutely puberulent. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 2.5–3 × 1–1.5 mm, lateral pair not saccate basally; petals white, oblanceolate, 6–8 × 2–3 mm (not clawed, apex obtuse to subemarginate); filaments: median pairs 3.5–4 mm, lateral pair 2–2.5 mm; anthers ovate, ca. 0.7 mm. |
sepals oblong, 5–7.5 × 1–2 mm, lateral pair slightly saccate basally; petals purple to pale pink, oblanceolate, 9–18 × 2–5 mm (clawed, apex rounded); filaments: median pairs 5–10 mm, lateral pair 3.5–8 mm; anthers linear, 1.5–3 mm. |
Fruiting pedicels | divaricate-ascending, (7–)10–17 mm. |
ascending to divaricate, 15–40 mm. |
Fruits | linear, (1.5–)2–3.5(–4) cm × 1.3–1.7 mm; ovules 10–16 per ovary; style 2–4 mm. |
linear, 2.5–4 cm × 1.5–2.5 mm; ovules 8–12 per ovary; style (5–)7–11 mm. |
Seeds | brown, oblong, 1.7–2 × 1–1.2 mm. |
dark brown, oblong, 2–2.5 × 1–1.5 mm. |
2n | = ca. 128. |
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Cardamine clematitis |
Cardamine angustata |
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Phenology | Flowering May–Jun. | Flowering Mar–May. |
Habitat | Wet areas, springs, moist slopes | Moist woods, wooded ridges and bottomlands, floodplains, shady ravines, streambeds |
Elevation | 1300-1800 m (4300-5900 ft) | 300-1300 m (1000-4300 ft) |
Distribution |
AL; FL; GA; NC; TN; VA |
AL; AR; DC; DE; GA; IN; KY; MD; MS; NC; NJ; OH; PA; SC; TN; VA; WV
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Discussion | Cardamine heterophylla (Nuttall) Alph. Wood (1870), not Host (1797) is an illegitimate name, sometimes found in synonymy with C. angustata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 470. | FNA vol. 7, p. 467. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. angustata var. ouachitana, Dentaria heterophylla | |
Name authority | Shuttleworth ex A. Gray: Proc. Amer. Acad. Arts 15: 45. (1880) | O. E. Schulz: Bot. Jahrb. Syst. 32: 349. (1903) |
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